Earlshall Muir

Site of Special Scientific Interest · 56.3906, -2.8391

Also within

Fife County

Condition (NatureScot)

Unfavourable Recovering (1) Unfavourable Declining (1) Favourable (3)
UnitFeatureConditionArea (ha)Assessed
F1 Breeding bird assemblage Unfavourable Recovering 2014-03-13
F2 Sand dunes Unfavourable Declining 2023-02-02
F3 Beetle assemblage Favourable 2015-05-20
F4 Grayling butterfly (Hipparchia semele) Favourable 2016-03-14
F5 Wet woodland Favourable 2012-11-01

Climate exposure

This is about exposure to a nationwide pressure this habitat type carries — not a verdict on this site's own management.

Coastal

Coastal habitats are exposed to sea-level rise and storm erosion. Where hard sea defences sit behind them, there's nowhere landward to retreat to — a squeeze that keeps compressing the habitat no matter how well it's managed.

Habitats

Dune slack, Sand dune

Geology & landscape

Landscape processes: Coastal

Birds & fish recorded

Birds

SpeciesSeason
lapwing Breeding
ringed plover Breeding
sedge warbler Breeding
shelduck Breeding
snipe Breeding
stonechat Breeding

Fish

SpeciesSeason
Grayling Unknown

Description

EARLSHALL MUIR Earlshall Muir Site of Special Scientific Interest (SSSI) is located approximately 2.5km east of Leuchars and 8km north of St. Andrews. The site represents the largest remaining tract of unafforested lime-poor sand dunes in south-east Scotland, with representative examples of dune ridge and dune slack plant communities. It is one of the few remaining examples in Scotland of a dune system exhibiting a complete succession of vegetation from accreting mobile dunes through to birch and alder woodland. The frontal dune system is in a dynamic state with natural processes continuing uninterrupted within the coastal cell leading to continuing accretion of the foreshore. Inland there is still some movement within the system with occasional blowouts occurring. There has been a large reduction in areas of young dune slack as conditions have become more stable. The site contains an extensive area of dune alder, willow and birch woodland which is now a rare habitat in Britain due to widespread reclamation and afforestation. A diverse breeding bird assemblage has been recorded including populations of shelduck Tadorna tadorna, snipe Gallinago gallinago, lapwing Vanellus vanellus, ringed plover Charadrius hiaticula, sedge warbler Acrocephalus schoenobaenus, reed bunting Emberiza schoeniclus and stonechat Saxicola torquatus. The invertebrate fauna is diverse including the grayling butterfly Hipparchia semele and several rare beetle species including Arena tabida and Pissodes validirostris. Although not notified as a feature, the site also supports a number of rare vascular plant species including the nationally rare large yellow sedge Carex flava, and the nationally scarce seaside centaury Centaurium littorale. Several locally rare plant species occur, including grass-of-Parnassus Parnassia palustris and field gentian Gentianella campestris.

Species mentioned

English name
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Latin name
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IUCN / Red List Type Family
stonechat Saxicola torquatus Not listed in GB list bird muscicapidae
reed bunting Emberiza schoeniclus LC (Breeding) bird-bunting emberizidae
shelduck Tadorna tadorna EN (NonBreeding), EN (Breeding) bird-duck anatidae
snipe Gallinago gallinago DD (NonBreeding) bird-snipe scolopacidae
ringed plover Charadrius hiaticula VU (NonBreeding) bird-wader charadriidae
lapwing Vanellus vanellus VU (NonBreeding), EN (Breeding) bird-wader charadriidae
sedge warbler Acrocephalus schoenobaenus LC (Breeding) bird-warbler acrocephalidae
grayling Hipparchia semele EN butterfly nymphalidae
seaside centuary Centaurium littorale LC herb gentianaceae
field gentian Gentianella campestris LC herb gentianaceae
grass of Parnassus Parnassia palustris VU herb celastraceae
yellow-green sedge Carex flava VU sedge cyperaceae

Plant communities

SD1 — Coastal vegetated shingle, Sand dune

Rumex crispus-Glaucium flavum shingle community

Solanum dulcamara, Crithmum maritimum, Cirsium arvense, Cirsium vulgare, Helminthotheca echioides, Hypochaeris radicata, Jacobaea vulgaris, Lactuca serriola, Leontodon saxatilis, Senecio viscosus, Sonchus arvensis, Sonchus asper, Tripleurospermum maritimum, Crambe maritima, Cerastium fontanum, Honckenya peploides, Sagina apetala, Silene uniflora, Atriplex prostrata, Sedum acre, Euphorbia paralias, Lathyrus japonicus, Geranium robertianum, Glaucium flavum, Plantago coronopus, Plantago lanceolata, Rumex crispus, Potentilla anserina, Ammophila arenaria, Arrhenatherum elatius, Elytrigia atherica, Elytrigia juncea, Festuca rubra, Holcus lanatus, Lolium perenne

SD2 — Coastal vegetated shingle, Sand dune

Honkenya peploides-Cakile maritima strandline community

Salsola kali, Achillea millefolium, Cirsium arvense, Eryngium maritimum, Hypochaeris radicata, Senecio vulgaris, Sonchus arvensis, Tripleurospermum maritimum, Cakile maritima, Cerastium diffusum, Honckenya peploides, Silene uniflora, Stellaria media agg., Atriplex glabriuscula, Atriplex laciniata, Atriplex littoralis, Atriplex patula, Atriplex prostrata, Sedum acre, Polygonum oxyspermum subsp. Raii, Rumex crispus, Potentilla anserina, Agrostis stolonifera, Ammophila arenaria, Arrhenatherum elatius, Elytrigia juncea, Elytrigia repens, Festuca rubra, Leymus arenarius, Poa annua

SD3 — Coastal vegetated shingle, Sand dune

Matricaria maritima-Galium aparine strandline community

Bolboschoenus maritimus, Juncus bufonius, Juncus gerardii, Ligusticum scoticum, Matricaria discoidea, Sonchus arvensis, Sonchus asper, Tripleurospermum maritimum, Mertensia maritima, Cakile maritima, Cochlearia officinalis, Honckenya peploides, Stellaria media agg., Atriplex glabriuscula, Atriplex patula, Atriplex prostrata, Triglochin maritimum, Plantago maritima, Armeria maritima, Rumex crispus, Glaux maritima, Galium aparine, Agrostis stolonifera, Elytrigia repens, Festuca rubra, Holcus lanatus, Leymus arenarius, Poa annua

SD4 — Sand dune

Elymus farctus ssp. boreali-atlanticus foredune community

Salsola kali, Artemisia maritima, Cirsium arvense, Cirsium vulgare, Eryngium maritimum, Hypochaeris radicata, Jacobaea vulgaris, Matricaria chamomilla, Senecio squalidus, Senecio vulgaris, Sonchus arvensis, Sonchus asper, Taraxacum officianalis, Tripleurospermum maritimum, Cakile maritima, Cerastium diffusum, Honckenya peploides, Atriplex glabriuscula, Atriplex laciniata, Atriplex prostrata, Euphorbia paralias, Polygonum oxyspermum subsp. Raii, Rumex crispus, Agrostis stolonifera, Ammophila arenaria, Elytrigia atherica, Elytrigia juncea, Festuca rubra, Leymus arenarius

SD5 — Sand dune

Leymus arenarius mobile dune community

Bryum algovicum, Ceratodon purpureus, Brachythecium rutabulum, Syntrichia ruraliformis, Cirsium arvense, Hypochaeris radicata, Sonchus arvensis, Sonchus asper, Taraxacum officianalis, Cakile maritima, Cerastium fontanum, Honckenya peploides, Atriplex glabriuscula, Atriplex prostrata, Chamerion angustifolium, Rumex crispus, Galium aparine, Ammophila arenaria, Elytrigia atherica, Elytrigia juncea, Elytrigia repens, Festuca rubra, Holcus lanatus, Leymus arenarius, Poa pratensis

SD6 — Sand dune

Ammophila arenaria mobile dune community

Lupinus arboreus, Carex arenaria, Ceratodon purpureus, Brachythecium albicans, Syntrichia ruraliformis, Angelica sylvestris, Heracleum sphondylium, Cirsium arvense, Cirsium vulgare, Eryngium maritimum, Hypochaeris radicata, Jacobaea vulgaris, Senecio squalidus, Senecio vulgaris, Sonchus arvensis, Sonchus asper, Taraxacum officianalis, Tussilago farfara, Cynoglossum officinale, Cakile maritima, Honckenya peploides, Atriplex glabriuscula, Atriplex laciniata, Calystegia soldanellla, Euphorbia paralias, Euphorbia portlandica, Trifolium arvense, Trifolium repens, Eriodium cicutarium, Chamerion angustifolium, Linaria vulgaris, Rumex crispus, Ranunculus repens, Rubus fruticosus agg., Galium verum, Urtica dioica, Agrostis capillaris, Agrostis stolonifera, Ammophila arenaria, Catapodium marinum, Dactylis glomerata, Elytrigia atherica, Elytrigia juncea, Elytrigia repens, Festuca rubra, Holcus lanatus, Leymus arenarius, Poa annua, Poa pratensis

SD7 — Sand dune

Ammophila arenaria-Festuca rubra semi-fixed dune community

Carex arenaria, Luzula campestris, Bryum argenteum, Bryum capillare, Ceratodon purpureus, Dicranum scoparium, Brachythecium albicans, Brachythecium rutabulum, Homalothecium lutescens, Hypnum cupressiforme, Kindbergia praelonga, Pseudoscleropodium purum, Rhytidiadelphus squarrosus, Rhytidiadelphus triquetrus, Syntrichia ruraliformis, Tortella flavovirens, Lophocolea bidentata, Cladonia fimbriata, Cladonia furcata, Cladonia impexa, Cladonia pyxidata, Cladonia rangiformis, Peltigera canina, Peltigera rufescens, Heracleum sphondylium, Achillea millefolium, Centaurea nigra, Cirsium arvense, Cirsium vulgare, Crepis capillaris, Eryngium maritimum, Hypochaeris radicata, Jacobaea vulgaris, Leontodon hispidus, Leontodon saxatilis, Pilosella officinarum, Scorzoneroides autumnalis, Senecio vulgaris, Sonchus arvensis, Sonchus asper, Taraxacum officianalis, Tragopogon pratensis, Tussilago farfara, Myosotis ramosissima, Erophila verna, Campanula rotundifolia, Valerianella locusta, Arenaria serpyllifolia, Cerastium diffusum, Cerastium fontanum, Cerastium semidecandrum, Calystegia soldanellla, Sedum acre, Euphorbia paralias, Euphorbia portlandica, Anthyllis vulneraria, Lotus corniculatus, Ononis repens, Trifolium arvense, Trifolium campestre, Trifolium repens, Vicia lathyroides, Vicia sativa, Centaurium erythraea, Eriodium cicutarium, Thymus praecox, Linum catharticum, Chamerion angustifolium, Rhinanthus minor, Plantago coronopus, Plantago lanceolata, Veronica arvensis, Veronica chamaedrys, Rumex crispus, Ranunculus bulbosus, Ranunculus repens, Acaena novae-zelandiae, Rubus caesius, Rubus fruticosus agg., Galium verum, Viola canina, Viola riviniana, Viola tricolor, Agrostis capillaris, Aira praecox, Ammophila arenaria, Arrhenatherum elatius, Dactylis glomerata, Elytrigia atherica, Elytrigia juncea, Elytrigia repens, Festuca rubra, Holcus lanatus, Leymus arenarius, Phleum arenarium, Poa pratensis

SD8 — Machair sand dune, Sand dune

Festuca rubra-Galium verum fixed dune grassland

Salix repens, Rosa spinosissima, Carex arenaria, Carex caryophyllea, Carex flacca, Luzula campestris, Plagiomnium rostratum, Plagiomnium undulatum, Ditrichum flexicaule, Brachythecium albicans, Calliergonell cuspidatum, Climacium dendroides, Entodon concinnus, Homalothecium lutescens, Hylocomium splendens, Hypnum cupressiforme, Kindbergia praelonga, Pleurozium schreberi, Pseudoscleropodium purum, Rhytidiadelphus squarrosus, Rhytidiadelphus triquetrus, Thuidium delicatulum, Thuidium tamariscinum, Syntrichia ruraliformis, Lophocolea bidentata, Cladonia rangiformis, Odontites verna, Peltigera canina, Peltigera rufescens, Angelica sylvestris, Daucus carota, Heracleum sphondylium, Achillea millefolium, Bellis perennis, Centaurea nigra, Cirsium arvense, Cirsium vulgare, Crepis capillaris, Hypochaeris radicata, Jacobaea vulgaris, Leucanthemum vulgare, Pilosella officinarum, Scorzoneroides autumnalis, Sonchus oleraceus, Taraxacum officianalis, Myosotis arvensis, Myosotis ramosissima, Campanula rotundifolia, Succisa pratensis, Cerastium arvense, Cerastium diffusum, Cerastium fontanum, Cerastium semidecandrum, Sedum acre, Equisetum arvense, Anthyllis vulneraria, Astragalus danicus, Lotus corniculatus, Medicago lupulina, Ononis repens, Trifolium campestre, Trifolium pratense, Trifolium repens, Vicia cracca, Vicia lathyroides, Gentianella amarella, Eriodium cicutarium, Thymus praecox, Linum catharticum, Dactylorhiza viridis, Neottia ovata, Euphrasia officinalis, Rhinanthus minor, Plantago coronopus, Plantago lanceolata, Plantago major, Plantago maritima, Veronica arvensis, Veronica chamaedrys, Polygala vulgaris, Rumex acetosa, Rumex acetosella, Primula veris, Primula vulgaris, Ranunculus acris, Ranunculus bulbosus, Ranunculus repens, Thalictrum minus, Potentilla anserina, Galium verum, Viola riviniana, Viola tricolor, Agrostis capillaris, Agrostis stolonifera, Aira praecox, Ammophila arenaria, Anthoxanthum odoratum, Arrhenatherum elatius, Avenula pubescens, Cynosurus cristatus, Dactylis glomerata, Elytrigia atherica, Elytrigia juncea, Elytrigia repens, Festuca ovina, Festuca rubra, Holcus lanatus, Koeleria macrantha, Leymus arenarius, Lolium perenne, Poa pratensis, Poa trivialis, Trisetum flavescens

SD9 — Sand dune

Ammophila arenaria-Arrhenatherum elatius dune grassland

Rosa spinosissima, Carex arenaria, Carex flacca, Luzula campestris, Brachythecium rutabulum, Hypnum cupressiforme, Pseudoscleropodium purum, Rhytidiadelphus squarrosus, Rhytidiadelphus triquetrus, Syntrichia ruraliformis, Heracleum sphondylium, Pimpinella saxifraga, Torilis japonica, Achillea millefolium, Bellis perennis, Centaurea nigra, Cirsium arvense, Crepis capillaris, Hypochaeris radicata, Jacobaea vulgaris, Leontodon hispidus, Pilosella officinarum, Taraxacum officianalis, Myosotis arvensis, Campanula rotundifolia, Cerastium fontanum, Silene latifolia, Equisetum arvense, Astragalus danicus, Lathyrus pratensis, Lotus corniculatus, Ononis repens, Trifolium repens, Vicia sativa, Geranium sanguineum, Thymus praecox, Plantago lanceolata, Veronica chamaedrys, Rumex acetosa, Primula veris, Ranunculus bulbosus, Thalictrum minus, Fragaria vesca, Potentilla reptans, Poterium sanguisorba, Cruciata laevipes, Galium verum, Viola riviniana, Agrostis capillaris, Ammophila arenaria, Anthoxanthum odoratum, Arrhenatherum elatius, Dactylis glomerata, Elytrigia atherica, Elytrigia juncea, Elytrigia repens, Festuca rubra, Holcus lanatus, Koeleria macrantha, Leymus arenarius, Poa pratensis, Trisetum flavescens, Pteridium aquilinum

SD10 — Sand dune

Carex arenaria dune community

Carex arenaria, Ceratodon purpureus, Hypnum cupressiforme, Achillea millefolium, Jacobaea vulgaris, Senecio vulgaris, Taraxacum officianalis, Tripleurospermum maritimum, Cerastium fontanum, Honckenya peploides, Astragalus danicus, Chamerion angustifolium, Plantago lanceolata, Rumex acetosella, Potentilla anserina, Galium verum, Agrostis capillaris, Aira praecox, Ammophila arenaria, Dactylis glomerata, Elytrigia juncea, Festuca ovina, Festuca rubra, Holcus lanatus, Koeleria macrantha

SD11 — Sand dune

Carex arenaria-Cornicularia aculeata dune community

Calluna vulgaris, Carex arenaria, Luzula campestris, Ceratodon purpureus, Dicranum scoparium, Racomitrium canescens sens.lat., Brachythecium albicans, Hypnum cupressiforme, Rhytidium rugosum, Polytrichum juniperinum, Polytrichum piliferum, Syntrichia ruraliformis, Ptilidium ciliare, Hypogymnia physodes, Cladonia arbuscula, Cladonia coccifera, Cladonia fimbriata, Cladonia foliacea, Cladonia furcata, Cladonia gracilis, Cladonia impexa, Cladonia pyxidata, Cladonia rangiferina, Cladonia rangiformis, Cladonia squamosa, Cladonia tenuis, Cladonia uncialis, Cornicularia aculeata, Hypochaeris radicata, Jacobaea vulgaris, Logfia minima, Teesdalia nudicaulis, Campanula rotundifolia, Sedum acre, Astragalus danicus, Eriodium cicutarium, Thymus praecox, Plantago lanceolata, Rumex acetosella, Galium saxatile, Galium verum, Viola canina, Viola tricolor, Agrostis capillaris, Aira praecox, Ammophila arenaria, Anthoxanthum odoratum, Corynephorus canescens, Festuca ovina, Festuca rubra, Phleum arenarium

SD12 — Sand dune

Carex arenaria-Festuca ovina-Agrostis capillaris dune grassland

Calluna vulgaris, Erica cinerea, Carex arenaria, Carex pilulifera, Luzula campestris, Ceratodon purpureus, Dicranum scoparium, Brachythecium albicans, Hylocomium splendens, Hypnum cupressiforme, Pleurozium schreberi, Pseudoscleropodium purum, Rhytidiadelphus squarrosus, Rhytidiadelphus triquetrus, Polytrichum juniperinum, Cladonia arbuscula, Cladonia furcata, Peltigera canina, Achillea millefolium, Cirsium arvense, Hypochaeris radicata, Jacobaea vulgaris, Pilosella officinarum, Campanula rotundifolia, Cerastium diffusum, Cerastium fontanum, Astragalus danicus, Lathyrus pratensis, Lotus corniculatus, Ononis repens, Trifolium repens, Vicia lathyroides, Geranium molle, Thymus praecox, Plantago lanceolata, Veronica officinalis, Rumex acetosa, Rumex acetosella, Primula vulgaris, Ranunculus repens, Potentilla erecta, Galium saxatile, Galium verum, Urtica dioica, Viola riviniana, Agrostis capillaris, Ammophila arenaria, Anthoxanthum odoratum, Danthonia decumbens, Deschampsia flexuosa, Festuca ovina, Festuca rubra, Holcus lanatus, Koeleria macrantha, Poa pratensis

SD13 — Dune slack, Sand dune

Sagina nodosa-Bryum pseudotriquetrum dune-slack community

Salix repens, Hippophae rhamnoides, Carex arenaria, Carex flacca, Carex serotina, Juncus articulatus, Bryum pseudotriquetrum, Amblystegium serpens, Didymodon tophaceus, Moerckia hibernica, Aneura pinguis, Pellia endiviifolia, Petalophyllum ralfsii, Hydrocotyle vulgaris, Jacobaea vulgaris, Leontodon hispidus, Pilosella officinarum, Pulicaria dysenterica, Scorzoneroides autumnalis, Sonchus arvensis, Taraxacum officianalis, Arenaria serpyllifolia, Cerastium fontanum, Sagina nodosa, Equisetum palustre, Equisetum variegatum, Pyrola rotundifolia, Anthyllis vulneraria, Lotus corniculatus, Ononis repens, Blackstonia perfoliata, Centaurium erythraea, Mentha aquatica, Epilobium palustre, Dactylorhiza incarnata, Epipactis palustris, Euphrasia officinalis, Polygala vulgaris, Primula vulgaris, Samolus valerandi, Rubus caesius, Galium verum, Agrostis stolonifera, Ammophila arenaria, Festuca rubra, Holcus lanatus, Phragmites australis, Poa annua, Poa pratensis

SD14 — Dune slack, Sand dune

Salix repens-Campylium stellatum dune-slack community

Salix caprea, Salix repens, Hippophae rhamnoides, Carex arenaria, Carex caryophyllea, Carex demissa, Carex flacca, Carex hirta, Carex hostiana, Carex nigra, Carex panicea, Carex serotina, Cladium mariscus, Eleocharis palustris, Eleocharis quinqueflora, Juncus acutus, Juncus articulatus, Juncus gerardii, Juncus inflexus, Juncus maritimus, Bryum pseudotriquetrum, Brachythecium rutabulum, Calliergonell cuspidatum, Drepanocladus aduncus, Drepanocladus sendtneri, Kindbergia praelonga, Pseudocalliergon lycopodioides, Pseudoscleropodium purum, Leiocolea badensis, Lophocolea bidentata, Moerckia hibernica, Preissia quadrata, Aneura pinguis, Pellia endiviifolia, Riccardia chamedryfolia, Angelica sylvestris, Oenanthe lachenalii, Hydrocotyle vulgaris, Bellis perennis, Eupatorium cannabinum, Jacobaea erucifolia, Jacobaea vulgaris, Leontodon hispidus, Leontodon saxatilis, Pulicaria dysenterica, Scorzoneroides autumnalis, Sonchus arvensis, Taraxacum officianalis, Succisa pratensis, Sagina nodosa, Parnassia palustris, Equisetum arvense, Equisetum fluviatile, Equisetum palustre, Equisetum variegatum, Pyrola rotundifolia, Anthyllis vulneraria, Lotus corniculatus, Lotus pedunculatus, Trifolium fragiferum, Trifolium pratense, Trifolium repens, Blackstonia perfoliata, Centaurium pulchellum, Gentianella amarella, Lycopus europaeus, Mentha aquatica, Linum catharticum, Epilobium parviflorum, Dactylorhiza incarnata, Dactylorhiza praetermissa, Dactylorhiza purpurella, Epipactis palustris, Gymnadenia conopsea, Liparis loeselii, Euphrasia officinalis, Rhinanthus minor, Plantago coronopus, Plantago lanceolata, Plantago major, Polygala vulgaris, Anagallis tenella, Glaux maritima, Primula vulgaris, Samolus valerandi, Ranunculus acris, Ranunculus flammula, Ranunculus repens, Fragaria vesca, Potentilla anserina, Rubus caesius, Galium palustre, Viola riviniana, Agrostis stolonifera, Anthoxanthum odoratum, Briza media, Danthonia decumbens, Festuca rubra, Holcus lanatus, Phragmites australis, Poa pratensis, Ophioglossum vulgatum

SD15 — Dune slack, Sand dune

Salix repens-Calliergon cuspidatum dune-slack community

Salix caprea, Salix repens, Solanum dulcamara, Bolboschoenus maritimus, Carex arenaria, Carex flacca, Carex hirta, Carex nigra, Carex panicea, Cladium mariscus, Eriophorum angustifolium, Eleocharis palustris, Juncus acutus, Juncus articulatus, Juncus inflexus, Juncus maritimus, Juncus subnodulosus, Brachythecium mildeanum, Calliergonell cuspidatum, Drepanocladus sendtneri, Homalothecium lutescens, Kindbergia praelonga, Pseudocalliergon lycopodioides, Pseudoscleropodium purum, Lophocolea bidentata, Angelica sylvestris, Oenanthe lachenalii, Hydrocotyle vulgaris, Caltha palustris, Centaurea nigra, Cirsium arvense, Cirsium palustre, Eupatorium cannabinum, Jacobaea vulgaris, Pulicaria dysenterica, Scorzoneroides autumnalis, Cardamine pratensis, Succisa pratensis, Silene flos-cuculi, Parnassia palustris, Equisetum arvense, Equisetum fluviatile, Equisetum palustre, Equisetum variegatum, Lathyrus pratensis, Lotus corniculatus, Lotus pedunculatus, Trifolium pratense, Trifolium repens, Vicia cracca, Vicia sativa, Iris pseudacorus, Glechoma hederacea, Lycopus europaeus, Mentha aquatica, Linum catharticum, Lythrum salicaria, Epilobium palustre, Epilobium parviflorum, Dactylorhiza incarnata, Dactylorhiza praetermissa, Epipactis palustris, Plantago lanceolata, Anagallis tenella, Glaux maritima, Lysimachia vulgaris, Primula vulgaris, Samolus valerandi, Ranunculus acris, Ranunculus flammula, Ranunculus repens, Agrimonia eupatoria, Filipendula ulmaria, Potentilla anserina, Potentilla reptans, Rubus caesius, Galium palustre, Scutellaria galericulata, Agrostis canina, Agrostis capillaris, Agrostis stolonifera, Arrhenatherum elatius, Danthonia decumbens, Festuca rubra, Holcus lanatus, Molinia caerulea, Phragmites australis, Poa pratensis, Ophioglossum vulgatum

SD16 — Dune slack, Sand dune

Salix repens-Holcus lanatus dune-slack community

Salix caprea, Salix repens, Betula pubescens, Quercus robur, Carex arenaria, Carex flacca, Carex nigra, Carex serotina, Eleocharis quinqueflora, Juncus acutus, Juncus articulatus, Juncus inflexus, Juncus maritimus, Luzula campestris, Bryum pseudotriquetrum, Amblystegium serpens, Brachythecium albicans, Brachythecium rutabulum, Calliergonell cuspidatum, Hypnum cupressiforme, Kindbergia praelonga, Pseudoscleropodium purum, Syntrichia ruraliformis, Lophocolea bidentata, Moerckia hibernica, Pellia endiviifolia, Petalophyllum ralfsii, Riccardia chamedryfolia, Hydrocotyle vulgaris, Bellis perennis, Carlina vulgaris, Cirsium arvense, Crepis capillaris, Erigeron acris, Eupatorium cannabinum, Hypochaeris radicata, Jacobaea vulgaris, Leontodon hispidus, Leontodon saxatilis, Pilosella officinarum, Pulicaria dysenterica, Scorzoneroides autumnalis, Sonchus arvensis, Taraxacum officianalis, Cerastium fontanum, Sagina nodosa, Parnassia palustris, Equisetum arvense, Equisetum palustre, Equisetum variegatum, Pyrola rotundifolia, Anthyllis vulneraria, Lotus corniculatus, Ononis repens, Trifolium dubium, Trifolium pratense, Trifolium repens, Gentianella amarella, Mentha aquatica, Thymus praecox, Linum catharticum, Epilobium palustre, Dactylorhiza incarnata, Epipactis helleborine, Epipactis palustris, Euphrasia officinalis, Rhinanthus minor, Plantago lanceolata, Anagallis tenella, Primula vulgaris, Samolus valerandi, Ranunculus acris, Ranunculus flammula, Ranunculus repens, Fragaria vesca, Potentilla anserina, Rubus caesius, Galium verum, Viola riviniana, Agrostis capillaris, Agrostis stolonifera, Ammophila arenaria, Cynosurus cristatus, Festuca rubra, Holcus lanatus, Phragmites australis, Poa annua, Poa pratensis

SD17 — Dune slack, Sand dune

Potentilla anserina-Carex nigra dune-slack community

Salix caprea, Salix repens, Carex arenaria, Carex disticha, Carex flacca, Carex nigra, Carex panicea, Eriophorum angustifolium, Eleocharis palustris, Juncus acutiflorus, Juncus articulatus, Juncus bufonius, Juncus effusus, Juncus inflexus, Plagiomnium rostratum, Brachythecium rutabulum, Calliergonell cuspidatum, Cratoneuron filicinum, Drepanocladus aduncus, Drepanocladus sendtneri, Homalothecium lutescens, Kindbergia praelonga, Rhytidiadelphus squarrosus, Lophocolea bidentata, Odontites verna, Angelica sylvestris, Hydrocotyle vulgaris, Bellis perennis, Caltha palustris, Centaurea nigra, Scorzoneroides autumnalis, Taraxacum officianalis, Myosotis laxa, Cardamine pratensis, Cerastium fontanum, Sagina procumbens, Silene flos-cuculi, Parnassia palustris, Equisetum arvense, Equisetum fluviatile, Equisetum palustre, Equisetum variegatum, Lathyrus pratensis, Lotus corniculatus, Trifolium repens, Vicia cracca, Iris pseudacorus, Triglochin maritimum, Mentha aquatica, Linum catharticum, Epilobium palustre, Dactylorhiza fuchsii, Dactylorhiza incarnata, Dactylorhiza purpurella, Epipactis palustris, Euphrasia officinalis, Pedicularis palustris, Rhinanthus minor, Plantago lanceolata, Plantago maritima, Persicaria maculosa, Rumex acetosa, Rumex crispus, Glaux maritima, Primula vulgaris, Ranunculus flammula, Ranunculus repens, Comarum palustre, Filipendula ulmaria, Potentilla anserina, Rubus caesius, Galium palustre, Galium verum, Agrostis capillaris, Agrostis stolonifera, Alopecurus geniculatus, Anthoxanthum odoratum, Cynosurus cristatus, Elytrigia repens, Festuca rubra, Holcus lanatus, Lolium perenne, Poa annua, Poa pratensis, Poa trivialis

SD18 — Sand dune

Hippophae rhamnoides dune scrub

Salix repens, Sambucus nigra, Hippophae rhamnoides, Crataegus monogyna, Solanum dulcamara, Carex arenaria, Ceratodon purpureus, Brachythecium albicans, Brachythecium rutabulum, Kindbergia praelonga, Cladonia fimbriata, Cladonia furcata, Daucus carota, Heracleum sphondylium, Achillea millefolium, Cirsium arvense, Cirsium vulgare, Hypochaeris radicata, Jacobaea vulgaris, Leontodon saxatilis, Sonchus arvensis, Taraxacum officianalis, Cynoglossum officinale, Diplotaxis tenuifolia, Cerastium diffusum, Cerastium semidecandrum, Honckenya peploides, Sagina procumbens, Stellaria media agg., Calystegia soldanellla, Convolvulus arvensis, Lotus corniculatus, Ononis repens, Glechoma hederacea, Montia perfoliata, Chamerion angustifolium, Anacamptis pyramidalis, Plantago lanceolata, Rubus caesius, Rubus fruticosus agg., Galium aparine, Galium verum, Urtica dioica, Agrostis stolonifera, Aira praecox, Ammophila arenaria, Arrhenatherum elatius, Dactylis glomerata, Elytrigia atherica, Elytrigia juncea, Festuca rubra, Holcus lanatus, Leymus arenarius, Phleum arenarium, Phragmites australis, Poa pratensis, Poa trivialis

SD19 — Sand dune

Phleum arenarium-Arenaria serpyllifolia dune annual community

Carex arenaria, Luzula campestris, Brachythecium albicans, Homalothecium lutescens, Hypnum cupressiforme, Syntrichia ruraliformis, Tortella flavovirens, Tortella tortuosa, Peltigera canina, Cirsium arvense, Crepis capillaris, Hypochaeris glabra, Jacobaea vulgaris, Logfia minima, Erophila verna, Arenaria serpyllifolia, Cerastium diffusum, Cerastium semidecandrum, Sedum acre, Euphorbia paralias, Anthyllis vulneraria, Lotus corniculatus, Ononis repens, Trifolium arvense, Trifolium campestre, Trifolium dubium, Trifolium repens, Centaurium erythraea, Eriodium cicutarium, Geranium molle, Thymus praecox, Plantago coronopus, Plantago lanceolata, Galium verum, Viola tricolor, Aira caryophyllea, Aira praecox, Ammophila arenaria, Anthoxanthum odoratum, Catapodium marinum, Festuca rubra, Mibora minima, Phleum arenarium, Poa pratensis, Vulpia fasciculata