Castell y Waun a'i Barcdir / Chirk Castle and Parkland

Site of Special Scientific Interest · 52.9363, -3.0873

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Clwyd County BRYNIAU CLWYD A DYFFRYN DYFRDWY/CLWYDIAN RANGE AND DEE VALLEY Scenic Area Offa's Dyke Path National Trail

Habitats

Ancient woodland, Fast flowing river and stream, Wood pasture

Geology & landscape

Geological Feature: Cave / Cavern, Geological Fault · Geological Period: Carboniferous, Silurian · Rock Type: Glacial Till, Millstone Grit, Sandstone

Landscape processes: Glacial, Karst, Volcanic / Igneous

Description

Description Castell y Waun a’i Barcdir/Chirk Castle and Parkland is of special interest for being one of the best examples of ancient wood pasture and parkland in Wales, containing a large number and diverse species of veteran and ancient trees. The site is also of special interest for the important invertebrate species that these trees support. Also of special interest is the breeding roost of lesser horseshoe bats Rhinolophus hipposideros in the castle buildings. One area of grassland supports a diverse assemblage of grassland fungi which is of special interest and of national importance, including 15 species of waxcap Hygrocybe spp. Chirk Castle sits on elevated ground at 200m above sea level, north of the Ceiriog valley and south of the Dee valley, 2km west of the town of Chirk. The ground around it is gently rolling, rising to a ridge to the west, but dropping quite steeply to the south into the valley of the River Ceiriog at 94m above sea level. The castle was built in 1282 by Roger Mortimer and has been continually occupied since that time. The bedrock geology underlying the site is predominantly Carboniferous Cefn y fedw Sandstone (Millstone Grit), which was used to build much of the castle. The remainder of the site is underlain by Silurian mudstones on the western margin, Carboniferous limestones and sandstones in the southern part, and Coal Measures mudstones on the eastern margin. Geological structure has also had an important influence on the landscape around Chirk Castle. The presence of the Chirk Castle Fault accounts for the steep climb between Home Farm and the castle, whilst the steep slope south of the castle is partly due to vigorous downcutting by the river along another fault. Much of the bedrock is mantled by glacial till and glaciofluvial sands and gravels deposited during the last Ice Age. The history of the parkland is well known. By the time of an estate survey by Robert Eggerley in 1391-92, the park was timber fenced and contained 100 acres of woodland as deer park. During the Civil War, in the middle of the seventeenth century, Charles Myddelton declared for Charles I. In 1659, Parliament ordered the castle to be taken down and all the trees in the parkland to be felled. The castle was repaired and the parkland was re-planted during a restoration period in the 1660s. In 1675, Sir Thomas Myddelton extended the park to the south and the east to hold 500 deer. Within what are currently the open areas of the park, very little planting is earlier than the 1760s, with most of the oak planting in the park taking place between 1800 and 1840 as part of a re-landscaping project carried out by William Emes. The majority of the trees in the parkland today have a girth of between 3m and 5m and are large, fully mature trees of mostly English or pendunculate oak Quercus robur, with some ash Fraxinus excelsior, sessile oak Quercus petraea, and occasional examples of horse chestnut Aesculus hippocastanum and sycamore Acer pseudoplatanus. The parkland also contains hawthorn Crataegus monogyna, an important source of nectar for some invertebrates. The woodlands also contain these tree species, along with sweet chestnut Castanea sativa and beech Fagus sylvatica. The original line of Offa’s Dyke passes through the parkland, just to the west of the castle. Along the line of the dyke are a large number of truly ancient trees, mainly English or pendunculate oak. One ancient tree of particular historic interest is the great Oak at the “Gate of the Dead”, on the southern side of the site, adjacent to the B4500. This tree is thought to date from the reign of King Egbert in 802 and has a trunk of over 10 metres in circumference. The over-mature nature of the trees, together with the preponderance of standing and fallen dead and dying wood, is ideal habitat for invertebrate species. The parkland at Chirk Castle is of national importance for its community of saproxylic (dead wood) invertebrates, mostly beetles. These comprise specialised invertebrates which are extremely localised in occurrence and are intimately associated with sites supporting a continuum of decaying timber. The few localities where populations of these rare or scarce species persist are generally relict patches of pasture-woodland or parkland. The site supports an outstanding assemblage of beetle species associated with over-mature trees, totalling 174 different species recorded to date – more than at any other site in Wales. Several of these species are extremely rare in Wales or, in some cases, known only from Chirk Castle. They include the false darkling beetles Hypulus quercinus and Abdera quadrifasciata, the spectacular yellow and black longhorn beetle Saperda scalaris, and the mould beetle Batrisodes venustus. Chirk Castle also has a significant assemblage of fungus gnat species (Diptera, Mycetophilidae), many of which are likely to be associated with bracket fungi on veteran trees, although the ecology of most species is not well known. The stable courtyard, built in 1768-9, abuts the southern side of the castle and contains the stables and laundry wing. Lesser horseshoe bats use the roof voids of the western and southern ends of the laundry wing during the summer months as a maternity roost. The site is of national importance for lesser horseshoe bats. This species was once widespread across the southern half of the British Isles as far north as Yorkshire, however their population and geographical range has now contracted and is restricted to Wales, the west Midlands and the south west of England. The Welsh population is of international significance. On leaving the roost, the lesser horseshoe bats follow walls, hedges and other landscape features extending from the roost to access the adjacent mature broadleaved woodland, which provides extensive feeding habitat. Several night roosts exist in outlying buildings to the south and west of the castle. The bats are also thought to use some areas of mature pine woodland for feeding in wet weather during the summer months. It is assumed that the lesser horseshoe bats recorded hibernating in Bronygarth or Ceiriog Cave include animals using the laundry wing of the castle during the summer. The cave is 0.6 km directly south of the castle on the southern side of the River Ceiriog. The Castell y Waun a’i Barcdir/Chirk Castle and Parklands SSSI includes the flight lines and feeding areas of the lesser horseshoe bats from the maternity roost to the cave. Other bat species that have been recorded at Chirk Castle, its associated buildings, Home Farm and in the surrounding woodland include common pipistrelle Pipistrellus pipistrellus, soprano pipistrelle Pipistrellus pygmaeus, Natterer’s bat Myotis nattereri, Daubenton’s bat Myotis daubentonii, noctule Nyctalus noctula and the brown long-eared bat Plecotus auritus. Areas of unimproved grassland within Baddys’ Park contain a rich grassland fungal community that has at least 15 species of waxcap Hygrocybe fungi, including the yellow foot waxcap H. flavipes, earthy waxcap H. fornicata, glutinous waxcap H. glutinipes and oily waxcap H. quieta. Remarks: Part owned by the National Trust. The SSSI is within the proposed extension to the Clwydian Range AONB. Chirk Castle is listed on the Register of Historic Parks and Gardens in Wales. Chirk Castle is a Grade I Listed Building. Chirk Castle gates and sections of Offa’s Dyke are Scheduled Ancient Monuments.

Species mentioned

English name
search for this in other sites		 Latin name
search with Google		 IUCN / Red List Type Family
Daubenton's bat Myotis daubentonii LC bat vespertillonidae
Natterer's bat Myotis nattereri LC bat vespertillonidae
noctule bat Nyctalus noctula LC bat vespertillonidae
common pipstrelle bat Pipistrellus pipistrellus LC bat vespertillonidae
Soprano pipistrelle bat Pipistrellus pygmaeus LC bat vespertillonidae
brown long-eared bat Plecotus auritus LC bat vespertillonidae
lesser horseshoe bat Rhinolophus hipposideros LC bat vespertillonidae
beetle Abdera quadrifasciata LC beetle melandryidae
no common name Hypulus quercinus LC beetle melandryidae
beetle Saperda scalaris LC beetle longhorn cerambycidae
rove beetle Batrisodes venustus Not listed in GB list beetle rove staphylinidae
hawthorn Crataegus monogyna LC shrub rosaceae
sycamore Acer pseudoplatanus Not listed in GB list tree aceraceae
horse chestnut Aesculus hippocastanum Not listed in GB list tree hippocastanaceae
sweet chestnut Castanea sativa LC tree fagaceae
beech Fagus sylvatica LC tree fabiaceae
ash Fraxinus excelsior LC tree oleaceae
sessile or durmast oak Quercus petraea LC tree fagaceae
common oak Quercus robur LC tree fagaceae

Plant communities

A9 — Fast flowing river and stream, Marginal and inundation, Pond ditch and canal, Sluggish river and stream

Potamogeton natans community

Schoenoplectus lacustris, Juncus bulbosus, Sparganium angustifolium, Sparganium erectum, Sparganium minimum, Alisma plantago-aquatica, Lobelia dortmanna, Equisetum fluviatile, Myriophyllum alterniflorum, Elodea canadensis, Lemna minor, Lemna trisulca, Nymphaea alba, Callitriche platycarpa, Callitriche stagnalis, Littorella uniflora, Potamogeton natans, Ranunculus trichophyllus, Glyceria fluitans, Chara subspinosa

A14 — Fast flowing river and stream

Myriophyllum alterniflorum community

Isolepis fluitans, Juncus bulbosus, Fontinalis antipyretica, Lobelia dortmanna, Equisetum fluviatile, Myriophyllum alterniflorum, Lemna minor, Callitriche stagnalis, Littorella uniflora, Potamogeton filiformis, Potamogeton gramineus, Potamogeton natans, Ranunculus peltatus, Chara spp., Nitella hyalina

A17 — Chalk river and stream, Fast flowing river and stream

Ranunculus penicillatus ssp. pseudofluitans community

Berula erecta, Nasturtium officinale, Elodea canadensis, Lemna minor, Callitriche stagnalis, Veronica beccabunga, Polygonum amphibium, Polygonum hydropiper, Potamogeton crispus, Potamogeton pectinatus, Ranunculus penicillatus ssp. pseudofluitans

A18 — Fast flowing river and stream

Ranunculus fluitans community

Fontinalis antipyretica, Fontinalis squamosa var. curnowii, Myriophyllum alterniflorum, Myriophyllum spicatum, Elodea canadensis, Mentha aquatica, Lemna gibba, Lemna minor, Potamogeton perfoliatus, Ranunculus fluitans, Glyceria fluitans, Phalaris arundinacea

W10 — Atlantic oak wood, Lowland dry oak and birch wood, Lowland mixed oak and ash wood, Wood pasture

Quercus robur-Pteridium aquilinum-Rubus fruticosus woodland

Hedera helix, Acer pseudoplatanus, Ilex aquifolium, Betula pendula, Betula pubescens, Carpinus betulus, Corylus avellana, Fagus sylvatica, Castanea sativa, Quercus hybrid, Quercus petraea, Quercus robur, Tilia cordata, Tilia x europaea, Fraxinus excelsior, Larix decidua, Pinus nigra, Pinus sylvestris, Pseudotsuga menziesii, Malus sylvestris, Prunus avium, Sorbus aucuparia, Alnus glutinosa, Populus tremula, Taxus baccata, Ulmus glabra, Sambucus nigra, Viburnum lantana, Viburnum opulus, Rhododendron ponticum, Vaccinium myrtillus, Cytisus scoparius, Ulex europaeus, Ligustrum vulgare, Crataegus laevigata, Crataegus monogyna, Prunus spinosa, Carex sylvatica, Juncus effusus, Luzula multiflora, Luzula pilosa, Luzula sylvatica, Mnium hornum, Plagiomnium rostratum, Plagiomnium undulatum, Ceratodon purpureus, Dicranella heteromalla, Dicranoweisia cirrata, Dicranum scoparium, Amblystegium serpens, Brachythecium rutabulum, Eurhynchium striatum, Hypnum cupressiforme, Isothecium myosuroides, Kindbergia praelonga, Plagiothecium denticulatum, Plagiothecium undulatum, Pseudoscleropodium purum, Pseudotaxiphyllum elegans, Thuidium tamariscinum, Atrichum undulatum, Polytrichum formosum, Lophocolea bidentata, Pellia epiphylla, Narcissus pseudonarcissus, Aegopodium podagraria, Anthriscus sylvestris, Conopodium majus, Heracleum sphondylium, Sanicula europaea, Jacobaea vulgaris, Solidago virgaurea, Lonicera periclymenum, Silene dioica, Stellaria holostea, Stellaria media agg., Euphorbia amygdaloides, Mercurialis perennis, Corydalis claviculata, Geranium robertianum, Ajuga reptans, Glechoma hederacea, Lamiastrum galeobdolon, Stachys sylvatica, Teucrium scorodonia, Hyacinthoides non-scripta, Chamerion angustifolium, Circaea lutetiana, Oxalis acetosella, Digitalis purpurea, Veronica chamaedrys, Rumex acetosa, Rumex sanguineus, Lysimachia nemorum, Lysimachia nummularia, Primula vulgaris, Anemone nemorosa, Ficaria verna, Ranunculus repens, Fragaria vesca, Potentilla sterilis, Rosa canina, Rubus fruticosus agg., Rubus idaeus, Galium aparine, Galium odoratum, Galium saxatile, Melanpyrum pratense, Urtica dioica, Viola riviniana, Agrostis capillaris, Anthoxanthum odoratum, Arrhenatherum elatius, Brachypodium sylvaticum, Bromopsis ramosa, Dactylis glomerata, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Holcus lanatus, Holcus mollis, Melica uniflora, Milium effusum, Poa nemoralis, Poa trivialis, Blechnum spicant, Pteridium aquilinum, Dryopteris borreri, Dryopteris dilatata, Dryopteris filix-mas, Thelypteris limbosperma, Athyrium filix-femina

W11 — Altantic hazel wood, Atlantic oak wood, Caledonian forest, Upland birch wood, Wood pasture

Quercus petraea-Betula pubescens-Oxalis acetosella woodland

Acer campestre, Ilex aquifolium, Betula pendula, Betula pubescens, Corylus avellana, Fagus sylvatica, Quercus hybrid, Quercus petraea, Quercus robur, Fraxinus excelsior, Larix decidua, Sorbus aucuparia, Calluna vulgaris, Erica cinerea, Juniperus communis, Vaccinium myrtillus, Cytisus scoparius, Crataegus monogyna, Rubus saxatilis, Carex pilulifera, Luzula campestris, Luzula multiflora, Luzula pilosa, Luzula sylvatica, Mnium hornum, Plagiomnium affine, Plagiomnium undulatum, Dicranum majus, Dicranum scoparium, Cirriphyllum piliferum, Eurhynchium striatum, Hylocomium splendens, Hypnum cupressiforme, Isothecium myosuroides, Kindbergia praelonga, Loeskeobryum brevirostre, Plagiothecium denticulatum, Plagiothecium undulatum, Pleurozium schreberi, Pseudoscleropodium purum, Rhytidiadelphus loreus, Rhytidiadelphus squarrosus, Rhytidiadelphus triquetrus, Thuidium tamariscinum, Atrichum undulatum, Polytrichum commune, Polytrichum formosum, Polytrichum longisetum, Sphagnum capillifolium, Diplophyllum albicans, Lophocolea bidentata, Plagiochila asplenioides, Plagiochila spinulosa, Angelica sylvestris, Conopodium majus, Campanula rotundifolia, Lonicera periclymenum, Succisa pratensis, Cerastium fontanum, Stellaria holostea, Pyrola minor, Vaccinium vitis-idaea, Lathyrus montanus, Geranium robertianum, Hypericum pulchrum, Ajuga reptans, Teucrium scorodonia, Convallaria majalis, Hyacinthoides non-scripta, Oxalis acetosella, Digitalis purpurea, Veronica chamaedrys, Veronica officinalis, Rumex acetosa, Lysimachia nemorum, Primula vulgaris, Trientalis europaea, Anemone nemorosa, Ranunculus acris, Ranunculus repens, Potentilla erecta, Rubus fruticosus agg., Rubus idaeus, Galium aparine, Galium saxatile, Melanpyrum pratense, Viola riviniana, Agrostis capillaris, Agrostis vinealis, Anthoxanthum odoratum, Brachypodium sylvaticum, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Festuca rubra, Holcus lanatus, Holcus mollis, Poa pratensis, Poa trivialis, Blechnum spicant, Pteridium aquilinum, Dryopteris borreri, Dryopteris dilatata, Dryopteris filix-mas, Thelypteris limbosperma, Athyrium filix-femina

W14 — Beech wood, Wood pasture

Fagus sylvatica-Rubus fruticosus woodland

Hedera helix, Salix caprea, Acer pseudoplatanus, Ilex aquifolium, Betula pendula, Betula pubescens, Corylus avellana, Fagus sylvatica, Quercus robur, Fraxinus excelsior, Pinus sylvestris, Prunus avium, Sorbus aria, Sorbus aucuparia, Taxus baccata, Ulmus glabra, Sambucus nigra, Ruscus aculeatus, Ligustrum vulgare, Carex remota, Carex sylvatica, Luzula pilosa, Mnium hornum, Dicranella heteromalla, Dicranum scoparium, Fissidens taxifolius, Brachythecium rutabulum, Hypnum cupressiforme, Isothecium myosuroides, Kindbergia praelonga, Plagiothecium denticulatum, Plagiothecium undulatum, Pseudotaxiphyllum elegans, Thuidium tamariscinum, Atrichum undulatum, Polytrichum commune, Polytrichum formosum, Arum maculatum, Lonicera periclymenum, Moehringia trinervia, Stellaria media agg., Euphorbia amygdaloides, Mercurialis perennis, Hypericum pulchrum, Lamiastrum galeobdolon, Hyacinthoides non-scripta, Circaea lutetiana, Epipactis helleborine, Oxalis acetosella, Digitalis purpurea, Anemone nemorosa, Ranunculus repens, Geum urbanum, Rubus fruticosus agg., Rubus idaeus, Galium odoratum, Galium verum, Urtica dioica, Viola riviniana, Agrostis capillaris, Agrostis stolonifera, Brachypodium sylvaticum, Bromopsis ramosa, Dactylis glomerata, Deschampsia cespitosa, Deschampsia flexuosa, Holcus lanatus, Holcus mollis, Melica uniflora, Milium effusum, Poa nemoralis, Poa trivialis, Schedonorus giganteus, Pteridium aquilinum, Dryopteris dilatata, Dryopteris filix-mas

W15 — Beech wood, Wood pasture

Fagus sylvatica-Deschampsia flexuosa woodland

Hedera helix, Acer pseudoplatanus, Ilex aquifolium, Betula pendula, Betula pubescens, Fagus sylvatica, Quercus hybrid, Quercus petraea, Quercus robur, Pinus sylvestris, Sorbus aucuparia, Taxus baccata, Calluna vulgaris, Rhododendron ponticum, Vaccinium myrtillus, Ruscus aculeatus, Carex pilulifera, Luzula pilosa, Luzula sylvatica, Mnium hornum, Orthodontium lineare, Dicranella heteromalla, Dicranum scoparium, Leucobryum glaucum, Hylocomium splendens, Hypnum cupressiforme, Kindbergia praelonga, Plagiothecium undulatum, Pseudotaxiphyllum elegans, Rhytidiadelphus loreus, Thuidium tamariscinum, Polytrichum formosum, Tetraphis pellucida, Calypogeia fissa, Diplophyllum albicans, Gymnocolea inflata, Lepidozia reptans, Cladonia fimbriata, Cladonia squamules, Lonicera periclymenum, Hyacinthoides non-scripta, Chamerion angustifolium, Oxalis acetosella, Rubus fruticosus agg., Melanpyrum pratense, Agrostis capillaris, Deschampsia flexuosa, Holcus lanatus, Holcus mollis, Molinia caerulea, Poa nemoralis, Blechnum spicant, Pteridium aquilinum, Dryopteris dilatata

W16 — Atlantic oak wood, Lowland dry oak and birch wood, Wood pasture

Quercus spp.-Betula spp.-Deschampsia flexuosa woodland

Hedera helix, Acer pseudoplatanus, Ilex aquifolium, Betula pendula, Betula pubescens, Corylus avellana, Fagus sylvatica, Castanea sativa, Quercus hybrid, Quercus petraea, Quercus robur, Pinus sylvestris, Sorbus aria, Sorbus aucuparia, Populus tremula, Calluna vulgaris, Erica cinerea, Rhododendron ponticum, Vaccinium myrtillus, Ulex europaeus, Ulex gallii, Frangula alnus, Crataegus monogyna, Luzula sylvatica, Mnium hornum, Orthodontium lineare, Pohlia nutans, Campylopus flexuosus, Campylopus pyriformis, Dicranella heteromalla, Dicranodontium denudatum, Dicranoweisia cirrata, Dicranum fuscescens, Dicranum scoparium, Leucobryum glaucum, Hypnum cupressiforme, Hypnum jutlandicum, Isothecium myosuroides, Kindbergia praelonga, Plagiothecium denticulatum, Plagiothecium nemorale, Plagiothecium succulentum, Plagiothecium undulatum, Pleurozium schreberi, Pseudoscleropodium purum, Pseudotaxiphyllum elegans, Rhytidiadelphus loreus, Rhytidiadelphus triquetrus, Thuidium tamariscinum, Tetraphis pellucida, Barbilophozia attenuata, Barbilophozia floerkei, Calypogeia fissa, Calypogeia trichomanis, Cephalozia bicuspidata, Gymnocolea inflata, Lepidozia reptans, Lophocolea bidentata, Lophocolea heterophylla, Cladonia coccifera, Cladonia coniocraea, Cladonia fimbriata, Cladonia polydactyla, Cladonia squamosa, Solidago virgaurea, Lonicera periclymenum, Corydalis claviculata, Teucrium scorodonia, Hyacinthoides non-scripta, Chamerion angustifolium, Oxalis acetosella, Digitalis purpurea, Rumex acetosella, Potentilla erecta, Rubus fruticosus agg., Galium saxatile, Agrostis capillaris, Agrostis stolonifera, Anthoxanthum odoratum, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Festuca rubra, Holcus lanatus, Holcus mollis, Molinia caerulea, Blechnum spicant, Pteridium aquilinum, Dryopteris dilatata

W17 — Altantic hazel wood, Atlantic oak wood, Caledonian forest, Upland birch wood, Wood pasture

Quercus petraea-Betula pubescens-Dicranum majus woodland

Hedera helix, Salix caprea, Acer pseudoplatanus, Ilex aquifolium, Betula pendula, Betula pubescens, Corylus avellana, Fagus sylvatica, Quercus hybrid, Quercus petraea, Quercus robur, Fraxinus excelsior, Sorbus aucuparia, Calluna vulgaris, Erica cinerea, Erica tetralix, Juniperus communis, Rhododendron ponticum, Vaccinium myrtillus, Crataegus monogyna, Carex pilulifera, Luzula multiflora, Luzula pilosa, Luzula sylvatica, Andreaea rupestris, Mnium hornum, Rhizomnium punctatum, Campylopus flexuosus, Dicranella heteromalla, Dicranodontium denudatum, Dicranum fuscescens, Dicranum majus, Dicranum scoparium, Dicranum scottianum, Leucobryum glaucum, Ptychomitrium polyphyllum, Racomitrium fasciculare, Racomitrium heterostichum, Racomitrium lanuginosum, Eurhynchium striatum, Heterocladium heteropterum, Hylocomiastrum umbratum, Hylocomium armoricum, Hylocomium splendens, Hypnum callichroum, Hypnum cupressiforme, Hypnum jutlandicum, Isothecium myosuroides, Kindbergia praelonga, Loeskeobryum brevirostre, Plagiothecium denticulatum, Plagiothecium undulatum, Pleurozium schreberi, Pseudoscleropodium purum, Pseudotaxiphyllum elegans, Ptilium crista-castrensis, Rhytidiadelphus loreus, Rhytidiadelphus squarrosus, Rhytidiadelphus triquetrus, Sematophyllum demissum, Sematophyllum micans, Thuidium delicatulum, Thuidium tamariscinum, Polytrichum commune, Polytrichum formosum, Sphagnum capillifolium, Sphagnum fimbriatum, Sphagnum palustre, Sphagnum russowii, Sphagnum subnitens, Tetraphis pellucida, Adelanthus decipiens, Anastrepta orcadensis, Barbilophozia attenuata, Barbilophozia floerkei, Bazzania tricrenata, Bazzania trilobata, Blepharostoma trichophyllum, Calypogeia fissa, Calypogeia muelleriana, Cephalozia bicuspidata, Diplophyllum albicans, Frullania tamarisci, Fuscocephaloziopsis lunulifolia, Harpanthus scutatus, Jamesoniella autumnalis, Jungermannia gracillima, Lepidozia pearsonii, Lepidozia reptans, Lophocolea bidentata, Lophocolea heterophylla, Lophozia ventricosa, Marsupella emarginata, Microlejeunea ulicina, Mylia taylorii, Nowellia curvifolia, Plagiochila asplenioides, Plagiochila atlantica, Plagiochila killarniensis, Plagiochila porelloides, Plagiochila punctata, Plagiochila spinulosa, Ptilidium ciliare, Saccogyna viticulosa, Scapania gracilis, Scapania nemorea, Scapania umbrosa, Tritomaria exsectiformis, Tritomaria quinquedentata, Cladonia arbuscula, Cladonia chlorophaea, Cladonia digitata, Cladonia furcata, Cladonia impexa, Cladonia polydactyla, Cladonia squamosa, Cladonia squamules, Cladonia subcervicornis, Parmelia saxatilis, Sphaerophorus fragilis, Sphaerophorus globosus, Solidago virgaurea, Lonicera periclymenum, Succisa pratensis, Cornus suecica, Hypericum pulchrum, Teucrium scorodonia, Hyacinthoides non-scripta, Goodyera repens, Oxalis acetosella, Digitalis purpurea, Rumex acetosa, Lysimachia nemorum, Trientalis europaea, Anemone nemorosa, Potentilla erecta, Rubus fruticosus agg., Rubus idaeus, Galium saxatile, Melanpyrum pratense, Viola riviniana, Agrostis capillaris, Agrostis stolonifera, Agrostis vinealis, Anthoxanthum odoratum, Dactylis glomerata, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Holcus lanatus, Holcus mollis, Molinia caerulea, Poa nemoralis, Poa pratensis, Blechnum spicant, Pteridium aquilinum, Dryopteris aemula, Dryopteris borreri, Dryopteris dilatata, Dryopteris filix-mas, Hymenophyllum tunbrigense, Hymenophyllum wilsonii, Polypodium vulgare, Thelypteris limbosperma, Athyrium filix-femina