Tweed Catchment Rivers - England: Till Catchment

Site of Special Scientific Interest · 55.5912, -2.0803

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Also within

Northumberland CountyRoxburgh, Ettrick and Lauderdale County NORTHUMBERLAND National Park

Condition (Natural England)

Unfavourable Recovering (4) Unfavourable No Change (4) Unfavourable Declining (1)
UnitFeatureConditionArea (ha)Assessed
NT898282 — Unfavourable Recovering 9.32 2010-12-17
NT903420 — Unfavourable No Change 81.68 2010-12-17
NT906305 — Unfavourable No Change 35.48 2010-12-17
NT989244 — Unfavourable Declining 15.17 2010-12-17
NU015234 — Unfavourable Recovering 4.85 2010-12-17
NU032297 — Unfavourable No Change 44.34 2010-12-17
NU033167 — Unfavourable Recovering 23.66 2013-03-25
NU046237 — Unfavourable Recovering 4.85 2010-12-17
NU055219 — Unfavourable No Change 44.34 2010-12-17

Habitats

Fast flowing river and stream, Mixed scrub, Montane scrub, Purple moor grass and rush pasture

Geology & landscape

Rock Type: Alluvium

Landscape processes: Fluvial, Glacial, Volcanic / Igneous

Birds & fish recorded

Birds

SpeciesSeason
goldeneye Breeding
sand martin Breeding

Fish

SpeciesSeason
Atlantic salmon Spawning
Atlantic salmon Unknown
European eel Spawning
Grayling Spawning
Pike Spawning
River lamprey Spawning
River lamprey Unknown
Sea lamprey Spawning
Sea trout Spawning

Description

Description As part of the whole River Tweed system, the Till Catchment Rivers are clean rivers of high conservation and ecological value. The vegetation types show a natural succession from mineral poor upland streams through to communities which are typical of mineral rich lowland rivers. Floating beds of water crowfoot, Ranunculus, are of international significance and the blooming of a diatom Didymosphenia in the headwaters draining the Cheviot is unique in England. The fish fauna is particularly significant, the area supports one of the most important game fisheries in England, with large migrations of salmon and also supports the three British species of lamprey. The Till catchment also contains important habitat for otters. General Description The Till Catchment Rivers drain the Cheviot Massif in all directions and join the River Tweed via the confluence with the River Till. The Bowmont Water and River Glen drain west and north and the Rivers Breamish and Till drain south and east. Minor tributaries include College Burn, Wooler Water, Harthope Burn and Lilburn Burn. Associated habitats include semi natural woodland, rough grassland and semi aquatic areas, found along the edges of the rivers. These habitats provide food and shelter for many species. The tributaries which drain the resistant volcanic rocks of the Cheviot Hills River Breamish, College Burn and Harthope Burn are typically fast flowing and sometimes torrential. As the bedrock around the central massif changes to cement stones, limestones and sandstones the river gradient becomes more gradual, with riffles, pools and waterfalls evident. In the flat flood plains around the foothills the rivers cut through the thick glacial drift deposits of alluvium, boulder clays, sands and gravels which overlay the bedrock. In the broad flood plains of the Milfield Plain the Rivers Till and Glen meander through these glacial deposits, where the channel course changes over time, leaving oxbow lakes, pools and marshy areas. The variation in mineral content of the catchment is reflected in the changing communities of aquatic plants. The plant species in the upper reaches, particularly those found higher than 200 metres above sea level, are typical of base and nutrient poor upland rivers. Moss species are common on the boulders and bedrock and include Fontinalis antipyretica, Hygroamblystegium fluviatile, Philonotis fontana, Rhyncostegium riparioides and Racomitrium aciculare. Filamentous green algae are also characteristic in the open, grazed moorland landscape. Other plant species are less abundant, but occasionally water starworts Callitriche spp. and water crowfoot species such as stream water crowfoot Ranunculus penicillatus, occur in slacks and riffles. A diatom, Didymosphenia, found in streams on the volcanic rocks in the College Valley, produces seasonal blooms, the Cheviots is the only place within England where this phenomenon is known to occur. Along the margins of the river channels, species such as water mint Mentha aquatica, reed canary grass Phalaris arundinacea, floating sweet grass Glyceria fluitans, procumbent pearlwort Sagina procumbens and blinks Montia fontana occur in places. The banks support species which are characteristic of an upland grassland environment and include purple moor grass Molinia caerulea, mat grass Nardus stricta, rushes Juncus spp. and scrub species such as gorse Ulex europaeus, hawthorn Crataegus monogyna and willows Salix spp. Alder Alnus glutinosa woodland occurs along stretches of the riverside. In the lower reaches the aquatic plants are typical of base and nutrient rich lowland rivers. Mosses and other lower plant species become increasingly scarce, but higher plants such as pondweeds Potamogeton spp. and water crowfoots Ranunculus spp. increase in abundance. Characteristic species of the larger river channels include river water crowfoot Ranunculus fluitans and stream water crowfoot Ranunculus penicillatus. The presence of curled and perfoliate pondweeds Potamogeton crispus and Potamogeton perfoliatus in the watercourse and branched bur reed Sparganium erectum on the river margins, indicate the base rich nature of the rivers. Along the banks of the lower Till, non native species such as giant hogweed, Heracleum mantegazzianum, Japanese knotweed Reynoutria japonica and Himalayan balsam Impatiens glandulifera have become established. Semi natural woodland occurs more frequently along the banks of the lower rivers, particularly along the Rivers Breamish and Till. Although some of these woodlands have been managed and contain non native species, they provide important habitat and cover for many bird and insect species. The fish fauna of this river system is particularly diverse and the Atlantic salmon Salmo salar, sea trout Salmo trutta trutta and brown trout Salmo trutta are particularly significant. There are two main runs of salmon, one in the spring and a larger one in the autumn. Sea trout runs follow a similar pattern to that of the salmon. These salmonid species require the gravel beds of rivers to spawn. Trout and salmon juveniles are present in all tributaries, though adult sea trout tend to spawn mainly in the upper reaches and the River Glen holds one of the highest densities of salmon in the catchment. The presence of lampreys throughout the catchment is also important. Brook lampreys Lampetra planeri and river lampreys Lampetra fluviatilis have been recorded in the Rivers Till, Breamish, Glen and the Bowmont Water. Sea lamprey Petromyzon marinus are also known to occur in these rivers. These species prefer the lower gradient, fast flowing rivers with boulders as spawning grounds and the juveniles show preference for silty areas in slower flowing waters. Other species found throughout the catchment include eels Anguilla anguilla, stone loach Barbatula barbatula, minnows Phoxinus phoxinus, three spined stickleback Gasterosteus aculeatus, pike Esox lucius and grayling Thymallus thymallus. The freshwater shrimp Gammarus gammarus is present throughout the catchment indicating the low pollution content of the waters. The whole system supports a rich insect fauna associated with the riverine and adjacent habitats including a large number and diversity of species. Particularly abundant groups include mayflies, caddis flies and stoneflies which in both the larval and flying phases are an abundant food source for the salmon and trout. The lower and middle reaches of the River Till and its tributaries are good habitat for otters. Through improved habitat management the number of otters breeding in the catchment is increasing. Water vole are present in small isolated populations. The site also supports populations of birds which are typical of riverine and upland habitats. Mute swans are abundant in the lower reaches of the Till and Glen rivers and Wooler Water where they nest and feed. Other species that occur are heron, goldeneye, goosanders, sand martin and, in the upper reaches, oystercatchers, common sandpipers, dippers, grey and yellow wagtails. Other Information Parts of this site overlap SSSIs notified separately under the Wildlife and Countryside Act 1981 as amended, Till Riverbanks SSSI, Harthope Burn SSSI and College Valley Woodlands SSSI. It also abuts The Allers and Lilburn Valley Woodlands SSSI. Otters and water vole are protected, under Schedule 5 of the Wildlife and Countryside Act 1981 as amended. Floating vegetation of Ranunculus of plain and submountainous rivers is a habitat listed in Annex 1 of the EC Habitats and Species Directive 92/43/ EEC. Of the species associated with these catchment rivers, Annexes IIa, at maa and Va of the EC Habitats and Species Directive 92/43/ EEC and Schedules 2 and 3 of The Conservation Natural Habitats, etc Regulations 1994 list the following as specially protected, otters Annexes IIa, at maa and Schedule 2, salmon Annexes IIa, Va and Schedule 3, all three species of lamprey Annex IIa, river lamprey Schedule 3. This site is part of the River Tweed listed as a Grade 1 site in A Nature Conservation Review, D A Ratcliffe 1977.

Species mentioned

English name
search for this in other sites		 Latin name
search with Google		 IUCN / Red List Type Family
goldeneye Bucephala clangula VU (NonBreeding), VU (Breeding) bird-duck anatidae
sand martin Riparia riparia LC (Breeding) bird-martin hirundinidae
eel Anguilla anguilla CR fish anguillidae
stone loach Barbatula barbatula LC fish nemacheilidae
pike Esox lucius LC fish esocidae
three spined stickleback Gasterosteus aculeatus LC fish gasterosteidae
minnow Phoxinus phoxinus LC fish cyprinidae
Atlantic salmon Salmo salar EN fish salmonidae
brown trout Salmo trutta LC fish salmonidae
sea trout Salmo trutta trutta Not listed in GB list fish salmonidae
grayling Thymallus thymallus LC fish salmonidae
floating sweet grass Glyceria fluitans LC grass poaceae
purple moor grass Molinia caerulea LC grass poaceae
mat grass Nardus stricta LC grass poaceae
reed canary grass Phalaris arundinacea LC grass poaceae
giant hogweed Heracleum mantegazzianum Not listed in GB list herb apiaceae
Himmalayan balsam Impatiens glandulifera Not listed in GB list herb balsaminaceae
water mint Mentha aquatica LC herb lamiaceae
blinks Montia fontana LC herb montiaceae
curled pondweed Potamogeton crispus LC herb potamogetonaceae
perfoliate pondweed Potamogeton perfoliatus LC herb potamogetonaceae
water crowfoots Ranunculus fluitans LC herb ranunculaceae
stream water crowfoot Ranunculus penicillatus LC herb ranunculaceae
procumbent pearlwort Sagina procumbens LC herb carophyllaceae
river lamprey Lampetra fluviatilis LC lamprey petromyzontidae
brook lamprey Lampetra planeri LC lamprey petromyzontidae
sea lamprey Petromyzon marinus LC lamprey petromyzontidae
greayer water moss Fontinalis antipyretica LC moss hypnales
brookside feather moss Hygroamblystegium fluviatile LC moss hypnales
fountain apple moss Philonotis fontana LC moss bryales
yellow fringe moss Racomitrium aciculare LC moss grimmiales
long beaked water feather moss Rhyncostegium riparioides Not listed in GB list moss brachytheciaceae
branched bur reed Sparganium erectum LC reed typhaceae
hawthorn Crataegus monogyna LC shrub rosaceae
gorse Ulex europaeus LC shrub leguminosae
alder Alnus glutinosa LC tree salicaceae

Plant communities

A9 — Fast flowing river and stream, Marginal and inundation, Pond ditch and canal, Sluggish river and stream

Potamogeton natans community

Schoenoplectus lacustris, Juncus bulbosus, Sparganium angustifolium, Sparganium erectum, Sparganium minimum, Alisma plantago-aquatica, Lobelia dortmanna, Equisetum fluviatile, Myriophyllum alterniflorum, Elodea canadensis, Lemna minor, Lemna trisulca, Nymphaea alba, Callitriche platycarpa, Callitriche stagnalis, Littorella uniflora, Potamogeton natans, Ranunculus trichophyllus, Glyceria fluitans, Chara subspinosa

A14 — Fast flowing river and stream

Myriophyllum alterniflorum community

Isolepis fluitans, Juncus bulbosus, Fontinalis antipyretica, Lobelia dortmanna, Equisetum fluviatile, Myriophyllum alterniflorum, Lemna minor, Callitriche stagnalis, Littorella uniflora, Potamogeton filiformis, Potamogeton gramineus, Potamogeton natans, Ranunculus peltatus, Chara spp., Nitella hyalina

A17 — Chalk river and stream, Fast flowing river and stream

Ranunculus penicillatus ssp. pseudofluitans community

Berula erecta, Nasturtium officinale, Elodea canadensis, Lemna minor, Callitriche stagnalis, Veronica beccabunga, Polygonum amphibium, Polygonum hydropiper, Potamogeton crispus, Potamogeton pectinatus, Ranunculus penicillatus ssp. pseudofluitans

A18 — Fast flowing river and stream

Ranunculus fluitans community

Fontinalis antipyretica, Fontinalis squamosa var. curnowii, Myriophyllum alterniflorum, Myriophyllum spicatum, Elodea canadensis, Mentha aquatica, Lemna gibba, Lemna minor, Potamogeton perfoliatus, Ranunculus fluitans, Glyceria fluitans, Phalaris arundinacea

M22 — Coastal floodplain grazing marsh, Lowland fen, Purple moor grass and rush pasture

Juncus subnodulosus-Cirsium palustre fen-meadow

Salix cinerea, Solanum dulcamara, Carex acutiformis, Carex disticha, Carex elata, Carex flacca, Carex hirta, Carex lepidocarpa, Carex nigra, Carex panicea, Carex rostrata, Cladium mariscus, Eriophorum angustifolium, Eleocharis palustris, Eleocharis uniglumis, Juncus articulatus, Juncus effusus, Juncus inflexus, Juncus subnodulosus, Luzula multiflora, Plagiomnium affine, Plagiomnium rostratum, Plagiomnium undulatum, Rhizomnium punctatum, Brachythecium rutabulum, Calliergonell cuspidatum, Cratoneuron filicinum, Pseudoscleropodium purum, Lophocolea bidentata, Angelica sylvestris, Berula erecta, Oenanthe lachenalii, Thyselium palustre, Hydrocotyle vulgaris, Achillea ptarmica, Caltha palustris, Centaurea nigra, Cirsium arvense, Cirsium dissectum, Cirsium palustre, Eupatorium cannabinum, Pulicaria dysenterica, Myosotis laxa, Myosotis scorpioides, Cardamine pratensis, Succisa pratensis, Valeriana dioica, Valeriana officinalis, Cerastium fontanum, Silene flos-cuculi, Stellaria palustris, Calystegia sepium, Equisetum fluviatile, Equisetum palustre, Lathyrus pratensis, Lotus pedunculatus, Trifolium pratense, Trifolium repens, Vicia cracca, Hypericum tetrapterum, Iris pseudacorus, Triglochin palustre, Ajuga reptans, Lycopus europaeus, Mentha aquatica, Lemna minor, Lythrum salicaria, Menyanthes trifoliata, Epilobium hirsutum, Epilobium palustre, Epilobium parviflorum, Dactylorhiza fuchsii, Dactylorhiza incarnata, Dactylorhiza praetermissa, Epipactis palustris, Pedicularis palustris, Rhinanthus minor, Plantago lanceolata, Veronica beccabunga, Polygonum amphibium, Rumex acetosa, Rumex conglomeratus, Rumex sanguineus, Lysimachia vulgaris, Primula vulgaris, Ranunculus acris, Ranunculus flammula, Ranunculus repens, Thalictrum flavum, Comarum palustre, Filipendula ulmaria, Potentilla anserina, Potentilla erecta, Potentilla reptans, Galium palustre, Galium uliginosum, Scrophularia auriculata, Scutellaria galericulata, Agrostis stolonifera, Anthoxanthum odoratum, Arrhenatherum elatius, Briza media, Calamagrostis canescens, Cynosurus cristatus, Deschampsia cespitosa, Festuca rubra, Holcus lanatus, Molinia caerulea, Phalaris arundinacea, Phleum pratense, Phragmites australis, Poa trivialis, Schedonorus arundinaceus, Ophioglossum vulgatum, Thelypteris palustris

M23 — Coastal floodplain grazing marsh, Purple moor grass and rush pasture, Upland rush pasture

Juncus effusus/acutiflorus-Galium palustre rush-pasture

Carex demissa, Carex echinata, Carex nigra, Carex panicea, Juncus acutiflorus, Juncus articulatus, Juncus bulbosus, Juncus conglomeratus, Juncus effusus, Luzula campestris, Luzula multiflora, Aulacomnium palustre, Brachythecium rivulare, Brachythecium rutabulum, Calliergon cordifolium, Calliergonell cuspidatum, Kindbergia praelonga, Rhytidiadelphus squarrosus, Polytrichum commune, Calypogeia fissa, Lophocolea bidentata, Pellia epiphylla, Angelica sylvestris, Carum verticillatum, Hydrocotyle vulgaris, Achillea ptarmica, Caltha palustris, Cirsium palustre, Senecio aquaticus, Taraxacum officianalis, Myosotis laxa, Myosotis secunda, Cardamine flexuosa, Cardamine pratensis, Succisa pratensis, Valeriana officinalis, Cerastium fontanum, Silene flos-cuculi, Stellaria alsine, Stellaria graminea, Equisetum fluviatile, Equisetum palustre, Lathyrus pratensis, Lotus pedunculatus, Trifolium repens, Iris pseudacorus, Mentha aquatica, Scutellaria minor, Lythrum salicaria, Narthecium ossifragum, Epilobium obscurum, Epilobium palustre, Dactylorhiza maculata, Plantago lanceolata, Veronica scutellata, Polygonum hydropiper, Rumex acetosa, Primula vulgaris, Ranunculus acris, Ranunculus flammula, Ranunculus repens, Comarum palustre, Filipendula ulmaria, Potentilla anserina, Potentilla erecta, Galium palustre, Viola palustris, Agrostis canina, Agrostis capillaris, Agrostis stolonifera, Anthoxanthum odoratum, Cynosurus cristatus, Deschampsia cespitosa, Festuca rubra, Glyceria fluitans, Holcus lanatus, Molinia caerulea, Poa pratensis, Poa trivialis, Dryopteris expansa, Athyrium filix-femina

M24 — Coastal floodplain grazing marsh, Lowland fen, Purple moor grass and rush pasture

Molinia caerulea-Cirsium dissectum fen-meadow

Salix cinerea, Salix repens, Alnus glutinosa, Calluna vulgaris, Erica tetralix, Carex hostiana, Carex nigra, Carex panicea, Carex pulicaris, Cladium mariscus, Eriophorum angustifolium, Schoenus nigricans, Juncus acutiflorus, Juncus articulatus, Juncus conglomeratus, Juncus effusus, Juncus subnodulosus, Luzula multiflora, Aulacomnium palustre, Brachythecium rutabulum, Calliergonell cuspidatum, Hypnum cupressiforme, Kindbergia praelonga, Pseudoscleropodium purum, Thuidium tamariscinum, Sphagnum auriculatum, Sphagnum subnitens, Angelica sylvestris, Hydrocotyle vulgaris, Achillea ptarmica, Centaurea nigra, Cirsium arvense, Cirsium dissectum, Cirsium palustre, Eupatorium cannabinum, Leontodon saxatilis, Pulicaria dysenterica, Senecio aquaticus, Serratula tinctoria, Cardamine pratensis, Succisa pratensis, Valeriana dioica, Cerastium fontanum, Silene flos-cuculi, Drosera rotundifolia, Equisetum palustre, Lathyrus pratensis, Lotus pedunculatus, Trifolium pratense, Vicia cracca, Hypericum tetrapterum, Hypericum undulatum, Mentha aquatica, Scutellaria minor, Linum catharticum, Lythrum salicaria, Narthecium ossifragum, Dactylorhiza fuchsii, Dactylorhiza incarnata, Dactylorhiza maculata, Epipactis palustris, Gymnadenia conopsea, Pedicularis sylvatica, Plantago lanceolata, Polygala vulgaris, Rumex acetosa, Anagallis tenella, Primula vulgaris, Ranunculus acris, Ranunculus flammula, Filipendula ulmaria, Potentilla anserina, Potentilla erecta, Potentilla reptans, Rubus fruticosus agg., Galium palustre, Galium uliginosum, Galium verum, Viola palustris, Agrostis canina, Agrostis stolonifera, Anthoxanthum odoratum, Briza media, Danthonia decumbens, Deschampsia cespitosa, Festuca rubra, Holcus lanatus, Molinia caerulea, Phragmites australis

M25 — Blanket bog, Coastal floodplain grazing marsh, Lowland fen, Purple moor grass and rush pasture, Upland acid grassland

Molinia caerulea-Potentilla erecta mire

Salix cinerea, Calluna vulgaris, Erica tetralix, Myrica gale, Vaccinium oxycoccos, Ulex europaeus, Ulex gallii, Carex echinata, Carex flacca, Carex nigra, Carex panicea, Eriophorum angustifolium, Schoenus nigricans, Juncus acutiflorus, Juncus articulatus, Juncus effusus, Juncus subnodulosus, Luzula campestris, Luzula multiflora, Aulacomnium palustre, Pohlia nutans, Calliergon giganteum, Calliergonell cuspidatum, Hypnum jutlandicum, Kindbergia praelonga, Pleurozium schreberi, Pseudoscleropodium purum, Rhytidiadelphus squarrosus, Polytrichum commune, Sphagnum auriculatum, Sphagnum capillifolium, Sphagnum fallax, Sphagnum palustre, Sphagnum papillosum, Sphagnum subnitens, Calypogeia fissa, Cephalozia bicuspidata, Lophocolea bidentata, Lophozia ventricosa, Angelica sylvestris, Carum verticillatum, Hydrocotyle vulgaris, Achillea ptarmica, Centaurea nigra, Cirsium palustre, Eupatorium cannabinum, Pulicaria dysenterica, Serratula tinctoria, Cardamine pratensis, Lobelia urens, Succisa pratensis, Valeriana officinalis, Equisetum palustre, Lotus pedunculatus, Hypericum pulchrum, Hypericum tetrapterum, Mentha aquatica, Scutellaria minor, Menyanthes trifoliata, Narthecium ossifragum, Epilobium palustre, Dactylorhiza maculata, Rumex acetosa, Ranunculus acris, Ranunculus flammula, Filipendula ulmaria, Potentilla erecta, Rubus fruticosus agg., Galium palustre, Viola palustris, Agrostis canina, Agrostis capillaris, Agrostis curtisii, Agrostis stolonifera, Anthoxanthum odoratum, Danthonia decumbens, Deschampsia cespitosa, Festuca ovina, Festuca rubra, Holcus lanatus, Molinia caerulea, Nardus stricta, Phragmites australis, Dryopteris carthusiana, Osmunda regalis

M26 — Limestone pavement, Purple moor grass and rush pasture

Molinia caerulea-Crepis paludosa mire

Carex appropinquata, Carex disticha, Carex echinata, Carex flacca, Carex hostiana, Carex nigra, Carex panicea, Carex pulicaris, Juncus acutiflorus, Juncus articulatus, Juncus conglomeratus, Aulacomnium palustre, Plagiomnium undulatum, Calliergonell cuspidatum, Campylium elodes, Climacium dendroides, Ctenidium molluscum, Hylocomium splendens, Pseudoscleropodium purum, Rhytidiadelphus squarrosus, Thuidium tamariscinum, Lophocolea bidentata, Plagiochila asplenioides, Angelica sylvestris, Caltha palustris, Centaurea nigra, Cirsium palustre, Crepis paludosa, Leontodon hispidus, Serratula tinctoria, Succisa pratensis, Valeriana dioica, Silene flos-cuculi, Equisetum fluviatile, Equisetum palustre, Lathyrus pratensis, Trifolium repens, Menyanthes trifoliata, Rhinanthus minor, Plantago lanceolata, Rumex acetosa, Primula farinosa, Primula vulgaris, Anemone nemorosa, Ranunculus acris, Ranunculus auricomus, Trollius europaeus, Comarum palustre, Filipendula ulmaria, Geum rivale, Potentilla erecta, Sanguisorba officinalis, Galium palustre, Galium uliginosum, Agrostis stolonifera, Anthoxanthum odoratum, Briza media, Deschampsia cespitosa, Festuca ovina, Festuca rubra, Holcus lanatus, Molinia caerulea, Phragmites australis

W20 — Montane scrub

Salix lapponum-Luzula sylvatica scrub

Salix arbuscula, Salix cinerea, Salix lapponum, Salix myrsinites, Salix phylicifolia, Salix reticulata, Empetrum nigrum, Calluna vulgaris, Salix lanata, Vaccinium myrtillus, Vaccinium uliginosum, Carex atrata, Carex bigelowii, Carex binervis, Carex flacca, Luzula multiflora, Luzula sylvatica, Huperzia selago, Selaginella selaginoides, Aulacomnium palustre, Mnium hornum, Philonotis fontana, Plagiomnium undulatum, Rhizomnium punctatum, Dicranum bonjeanii, Dicranum majus, Dicranum scoparium, Fissidens adianthoides, Racomitrium lanuginosum, Calliergonell cuspidatum, Ctenidium molluscum, Hylocomium splendens, Hypnum callichroum, Hypnum cupressiforme, Palustriella commutatum, Plagiothecium denticulatum, Plagiothecium undulatum, Pleurozium schreberi, Ptilium crista-castrensis, Rhytidiadelphus loreus, Rhytidiadelphus squarrosus, Rhytidiadelphus triquetrus, Sanionia uncinata, Scorpidium revolvens, Thuidium tamariscinum, Atrichum undulatum, Polytrichastrum alpinum, Sphagnum capillifolium, Sphagnum fallax, Sphagnum girgensohnii, Sphagnum palustre, Sphagnum russowii, Sphagnum squarrosum, Sphagnum subnitens, Sphagnum teres, Barbilophozia barbata, Chiloscyphus polyanthos, Diplophyllum albicans, Lophocolea bidentata, Plagiochila porelloides, Ptilidium ciliare, Scapania undulata, Cladonia squamosa, Peltigera canina, Angelica sylvestris, Caltha palustris, Hieraclium spp., Saussurea alpina, Solidago virgaurea, Campanula rotundifolia, Succisa pratensis, Valeriana officinalis, Rhodiola rosea, Vaccinium vitis-idaea, Thymus praecox, Epilobium anagallidifolium, Dactylorhiza viridis, Euphrasia officinalis, Rhinanthus minor, Oxalis acetosella, Oxyria digyna, Polygonum viviparum, Rumex acetosa, Anemone nemorosa, Ranunculus acris, Thalictrum alpinum, Alchemilla alpina, Alchemilla glabra, Filipendula ulmaria, Geum rivale, Galium boreale, Galium saxatile, Saxifraga aizoides, Saxifraga hypnoides, Saxifraga oppositifolia, Saxifraga stellaris, Viola riviniana, Agrostis canina, Agrostis capillaris, Anthoxanthum odoratum, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Festuca rubra, Nardus stricta, Dryopteris dilatata, Phegopteris connectilis

W21 — Hedgerow scrub, Mixed scrub

Crataegus monogyna-Hedera helix scrub

Hedera helix, Salix cinerea, Acer pseudoplatanus, Ilex aquifolium, Betula pendula, Corylus avellana, Fagus sylvatica, Quercus robur, Fraxinus excelsior, Malus sylvestris, Sorbus aria, Taxus baccata, Ulmus glabra, Ulmus procera, Sambucus nigra, Viburnum lantana, Euonymus europaeus, Cornus sanguinea, Juniperus communis, Ligustrum vulgare, Clematis vitalba, Rhamnus cathartica, Crataegus laevigata, Crataegus monogyna, Prunus spinosa, Juncus effusus, Plagiomnium undulatum, Brachythecium rutabulum, Kindbergia praelonga, Rhynchostegium confertum, Tamus communis, Heracleum sphondylium, Torilis japonica, Arum maculatum, Arctium minus, Cirsium arvense, Lonicera periclymenum, Moehringia trinervia, Silene dioica, Stellaria graminea, Stellaria holostea, Stellaria media agg., Calystegia sepium, Mercurialis perennis, Geranium robertianum, Iris foetidissima, Glechoma hederacea, Lamium purpureum, Origanum vulgare, Stachys sylvatica, Teucrium scorodonia, Allium ursinum, Hyacinthoides non-scripta, Circaea lutetiana, Rumex sanguineus, Anemone nemorosa, Ranunculus repens, Fragaria vesca, Geum urbanum, Potentilla sterilis, Poterium sanguisorba, Rosa arvensis, Rosa canina, Rubia peregrina, Rubus fruticosus agg., Galium aparine, Urtica dioica, Viola riviniana, Anisantha sterilis, Arrhenatherum elatius, Brachypodium pinnatum, Brachypodium sylvaticum, Bromopsis erecta, Dactylis glomerata, Elytrigia repens, Holcus lanatus, Holcus mollis, Poa trivialis, Asplenium scolopendrium, Pteridium aquilinum, Dryopteris filix-mas

W22 — Hedgerow scrub, Mixed scrub

Prunus spinosa-Rubus fruticosus scrub

Hedera helix, Acer campestre, Corylus avellana, Ulex europaeus, Prunus spinosa, Plagiomnium undulatum, Brachythecium rutabulum, Eurhynchium striatum, Kindbergia praelonga, Thuidium tamariscinum, Atrichum undulatum, Lophocolea bidentata, Achillea millefolium, Cardamine flexuosa, Lonicera periclymenum, Moehringia trinervia, Silene dioica, Silene uniflora, Stellaria media agg., Mercurialis perennis, Geranium robertianum, Stachys sylvatica, Teucrium scorodonia, Hyacinthoides non-scripta, Oxalis acetosella, Digitalis purpurea, Plantago lanceolata, Veronica chamaedrys, Armeria maritima, Rumex acetosa, Filipendula ulmaria, Geum urbanum, Rosa canina, Rubus fruticosus agg., Rubus idaeus, Galium aparine, Urtica dioica, Viola riviniana, Agrostis capillaris, Brachypodium sylvaticum, Dactylis glomerata, Festuca rubra, Holcus lanatus, Holcus mollis, Poa trivialis, Asplenium scolopendrium, Pteridium aquilinum, Dryopteris borreri, Dryopteris filix-mas