Combe Wood and Linkenholt Hanging

Site of Special Scientific Interest · 51.3338, -1.4888

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Also within

Berkshire CountyHampshire County North Wessex Downs Scenic Area

Condition (Natural England)

Favourable (3)
UnitFeatureConditionArea (ha)Assessed
SU354597 — Favourable 6.68 2019-01-18
SU356597 — Favourable 51.98 2019-01-18
SU358592 — Favourable 47.82 2019-01-18

Habitats

Ancient woodland, Bracken scrub, Coppiced woodland, Mixed scrub, Montane scrub, Wood pasture

Geology & landscape

Rock Type: Chalk

Description

Area, 107.5 hectares 265.8 ac Description An extensive area of semi natural woodland situated in a sheltered valley near to the point where the counties of Berkshire, Hampshire and Wiltshire meet. The woodland is partly ancient and partly of secondary original, incorporating some small areas of chalk grassland and scrub which are relicts of open downland. The site lies predominantly on well drained, shallow rendzina soils overlying Upper Chalk; with deep calcareous loams occurring in the valley bottoms and more acidic soils derived from clay with flints on some plateau areas. The woodland consists of stands of ash field maple on the more calcareous soils and birch pedunculate oak on the acidic clays. Wych elm, beech and whitebeam occur locally. The northern part of the wood appears to have been managed as 'wood pasture' at some stage during its history, as indicated by the open structure of mature oaks, including some pollards and the sparse understorey of hazel with a very poor ground flora. Other parts of the wood consist of ancient coppice, the best example being Linkenholt Hanging which has some fine stands of ash and maple, a well developed hazel understorey and a diverse ground flora. The structural diversity of the wood, combined with the sheltered, humid situation provide ideal conditions for many species of mosses and lichens to grow, particularly epiphytic species which colonise the trunks and branches of trees. Typical of ancient woodland are the lichens Bacidia biatorina, Bacidia atropurpurea, Opegrapha ochrocheila, Pertusaria hemisphaerica and Thelotrema lepadinum and the mosses Orthotrichum lyellii and Zygodon rupestris. Rare lichens include Strangospora ochrophora which in Britain is rare outside western valleys and Peltigera Melanops horizontalis which occurs in old forests. Uncommon mosses include Leucodon sciuroides, Neckera pumila and a rare variety of Heterocladium heteropterum which was discovered growing on chalk pebbles in the wood. The ground flora is dominated by bracken Pteridium aquilinum and rosebay willowherb Chamerion angustifolium in more open parts or bluebell Hyacinthoides non-scripta and dog's mercury Mercurialis perennis in areas where the canopy is denser. Many species typical of ancient woodland also occur, including wood spurge Euphorbia amygdaloides, Solomon's seal Polygonatum multiflorum, wood anemone Anemone nemorosa and early purple orchid Orchis mascula. Uncommon species include wood forget-me-not Myosotis sylvatica, the parasitic toothwort Lathraea squamaria and wild daffodil Narcissus pseudonarcissus. Some species such as herb paris Paris quadrifolia and sweet woodruff Galium odoratum, grow on base rich soils and are locally frequent in some parts of the wood. A variety of woodland insects are known from the site, including many beetles Coleoptera and flies Diptera which are of ten associated with dead or decaying wood. Species recorded include the cardinal beetle Pyrochroa coccinea, the soldier beetles Malthodes fibulatus, M. maurus and M. mysticus, the hoverflies Volucella pellucens and Baccha sp. and the robber fly Neoitamus cyanurus. Butterflies include speckled wood, white admiral and purple hairstreak. The clearings and woodland edges also attract many species of insects, including the hoverfly Platycheirus scutatus. Birds are numerous and hole nesting species such as woodpecker, tits and nuthatch are well represented. A large herd of fallow deer is resident in the wood and exerts a powerful influence on regeneration. The remnants of grassy downland support a rich chalk flora which includes autumn gentian Gentianella amarella, meadow saxifrage Saxifraga granulataa, clustered bellflower Campanula glomerata and common spotted orchid Dactylorhiza fuchsii. Nearby areas of scrub are composed of hawthorn Crataegus monogyna, dogwood Cornus sanguinea, wild privet Ligustrum vulgare and other shrubs. These areas support many insects including the Pyralid moth Pyrausta nigrata, the common ground hopper Tetrix undulata and butterflies such as marbled white and dark green fritillary, a declining species in Britain. The day flying humming bird hawk moth Macroglossum stellatarum, normally a continental immigrant, has also been recorded.

Species mentioned

English name
search for this in other sites		 Latin name
search with Google		 IUCN / Red List Type Family
black-headed cardinal beetle Pyrochroa coccinea LC beetle cardinal pyrochroidae
nuthatch Sitta europaea LC (Breeding) bird-nuthatch sittidae
bracken Pteridium aquilinum LC fern dennstaedtiaceae
robber fly Neoitamus cyanurus LC fly asilidae
lichen species name Melanops horizontalis Not listed in GB list fungus botryosphaeriaceae
common groundhopper Tetrix undulata LC groundhoppers tetrigidae
wood anemone Anemone nemorosa LC herb ranunculaceae
clustered bellflower Campanula glomerata VU herb campanulaceae
rosebay willowherb Chamerion angustifolium Not listed in GB list herb onagraceae
spotted orchids Dactylorhiza fuchsii LC herb orchidaceae
wood spurge Euphorbia amygdaloides LC herb euphorbiaceae
woodruff Galium odoratum LC herb rubiaceae
autumn gentian Gentianella amarella LC herb gentianaceae
bluebell Hyacinthoides non-scripta LC herb liliaceae
toothwort Lathraea squamaria LC herb orobanchaceae
dog's mercury Mercurialis perennis LC herb euphorbiaceae
wood forget-me-nots Myosotis sylvatica LC herb boraginaceae
wild daffodil Narcissus pseudonarcissus LC herb amaryllidaceae
early purple orchid Orchis mascula LC herb orchidaceae
herb paris Paris quadrifolia VU herb melanthiaceae
Soloman's seal Polygonatum multiflorum LC herb liliaceae
lichen Bacidia biatorina LC lichen ramalinaceae
no common name Strangospora ochrophora Not listed in GB list lichen no common name
barnacle lichen Thelotrema lepadinum LC lichen no common name
wry leaved tamarisk moss Heterocladium heteropterum LC moss hypnales
squirrel tail moss Leucodon sciuroides LC moss hypnales
dwarf neckera Neckera pumila LC moss hypnales
Lyell's bristle moss Orthotrichum lyellii Not listed in GB list moss orthotrichales
park yoke moss Zygodon rupestris LC moss orthotrichales
hummingbird hawk-moth Macroglossum stellatarum Not listed in GB list moth sphingidae
no common name Pyrausta nigrata Not listed in GB list moth crambidae
dogwood Cornus sanguinea LC shrub cornaceae
hawthorn Crataegus monogyna LC shrub rosaceae
wild privet Ligustrum vulgare LC shrub oleaceae

Plant communities

W10 — Atlantic oak wood, Lowland dry oak and birch wood, Lowland mixed oak and ash wood, Wood pasture

Quercus robur-Pteridium aquilinum-Rubus fruticosus woodland

Hedera helix, Acer pseudoplatanus, Ilex aquifolium, Betula pendula, Betula pubescens, Carpinus betulus, Corylus avellana, Fagus sylvatica, Castanea sativa, Quercus hybrid, Quercus petraea, Quercus robur, Tilia cordata, Tilia x europaea, Fraxinus excelsior, Larix decidua, Pinus nigra, Pinus sylvestris, Pseudotsuga menziesii, Malus sylvestris, Prunus avium, Sorbus aucuparia, Alnus glutinosa, Populus tremula, Taxus baccata, Ulmus glabra, Sambucus nigra, Viburnum lantana, Viburnum opulus, Rhododendron ponticum, Vaccinium myrtillus, Cytisus scoparius, Ulex europaeus, Ligustrum vulgare, Crataegus laevigata, Crataegus monogyna, Prunus spinosa, Carex sylvatica, Juncus effusus, Luzula multiflora, Luzula pilosa, Luzula sylvatica, Mnium hornum, Plagiomnium rostratum, Plagiomnium undulatum, Ceratodon purpureus, Dicranella heteromalla, Dicranoweisia cirrata, Dicranum scoparium, Amblystegium serpens, Brachythecium rutabulum, Eurhynchium striatum, Hypnum cupressiforme, Isothecium myosuroides, Kindbergia praelonga, Plagiothecium denticulatum, Plagiothecium undulatum, Pseudoscleropodium purum, Pseudotaxiphyllum elegans, Thuidium tamariscinum, Atrichum undulatum, Polytrichum formosum, Lophocolea bidentata, Pellia epiphylla, Narcissus pseudonarcissus, Aegopodium podagraria, Anthriscus sylvestris, Conopodium majus, Heracleum sphondylium, Sanicula europaea, Jacobaea vulgaris, Solidago virgaurea, Lonicera periclymenum, Silene dioica, Stellaria holostea, Stellaria media agg., Euphorbia amygdaloides, Mercurialis perennis, Corydalis claviculata, Geranium robertianum, Ajuga reptans, Glechoma hederacea, Lamiastrum galeobdolon, Stachys sylvatica, Teucrium scorodonia, Hyacinthoides non-scripta, Chamerion angustifolium, Circaea lutetiana, Oxalis acetosella, Digitalis purpurea, Veronica chamaedrys, Rumex acetosa, Rumex sanguineus, Lysimachia nemorum, Lysimachia nummularia, Primula vulgaris, Anemone nemorosa, Ficaria verna, Ranunculus repens, Fragaria vesca, Potentilla sterilis, Rosa canina, Rubus fruticosus agg., Rubus idaeus, Galium aparine, Galium odoratum, Galium saxatile, Melanpyrum pratense, Urtica dioica, Viola riviniana, Agrostis capillaris, Anthoxanthum odoratum, Arrhenatherum elatius, Brachypodium sylvaticum, Bromopsis ramosa, Dactylis glomerata, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Holcus lanatus, Holcus mollis, Melica uniflora, Milium effusum, Poa nemoralis, Poa trivialis, Blechnum spicant, Pteridium aquilinum, Dryopteris borreri, Dryopteris dilatata, Dryopteris filix-mas, Thelypteris limbosperma, Athyrium filix-femina

W11 — Altantic hazel wood, Atlantic oak wood, Caledonian forest, Upland birch wood, Wood pasture

Quercus petraea-Betula pubescens-Oxalis acetosella woodland

Acer campestre, Ilex aquifolium, Betula pendula, Betula pubescens, Corylus avellana, Fagus sylvatica, Quercus hybrid, Quercus petraea, Quercus robur, Fraxinus excelsior, Larix decidua, Sorbus aucuparia, Calluna vulgaris, Erica cinerea, Juniperus communis, Vaccinium myrtillus, Cytisus scoparius, Crataegus monogyna, Rubus saxatilis, Carex pilulifera, Luzula campestris, Luzula multiflora, Luzula pilosa, Luzula sylvatica, Mnium hornum, Plagiomnium affine, Plagiomnium undulatum, Dicranum majus, Dicranum scoparium, Cirriphyllum piliferum, Eurhynchium striatum, Hylocomium splendens, Hypnum cupressiforme, Isothecium myosuroides, Kindbergia praelonga, Loeskeobryum brevirostre, Plagiothecium denticulatum, Plagiothecium undulatum, Pleurozium schreberi, Pseudoscleropodium purum, Rhytidiadelphus loreus, Rhytidiadelphus squarrosus, Rhytidiadelphus triquetrus, Thuidium tamariscinum, Atrichum undulatum, Polytrichum commune, Polytrichum formosum, Polytrichum longisetum, Sphagnum capillifolium, Diplophyllum albicans, Lophocolea bidentata, Plagiochila asplenioides, Plagiochila spinulosa, Angelica sylvestris, Conopodium majus, Campanula rotundifolia, Lonicera periclymenum, Succisa pratensis, Cerastium fontanum, Stellaria holostea, Pyrola minor, Vaccinium vitis-idaea, Lathyrus montanus, Geranium robertianum, Hypericum pulchrum, Ajuga reptans, Teucrium scorodonia, Convallaria majalis, Hyacinthoides non-scripta, Oxalis acetosella, Digitalis purpurea, Veronica chamaedrys, Veronica officinalis, Rumex acetosa, Lysimachia nemorum, Primula vulgaris, Trientalis europaea, Anemone nemorosa, Ranunculus acris, Ranunculus repens, Potentilla erecta, Rubus fruticosus agg., Rubus idaeus, Galium aparine, Galium saxatile, Melanpyrum pratense, Viola riviniana, Agrostis capillaris, Agrostis vinealis, Anthoxanthum odoratum, Brachypodium sylvaticum, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Festuca rubra, Holcus lanatus, Holcus mollis, Poa pratensis, Poa trivialis, Blechnum spicant, Pteridium aquilinum, Dryopteris borreri, Dryopteris dilatata, Dryopteris filix-mas, Thelypteris limbosperma, Athyrium filix-femina

W14 — Beech wood, Wood pasture

Fagus sylvatica-Rubus fruticosus woodland

Hedera helix, Salix caprea, Acer pseudoplatanus, Ilex aquifolium, Betula pendula, Betula pubescens, Corylus avellana, Fagus sylvatica, Quercus robur, Fraxinus excelsior, Pinus sylvestris, Prunus avium, Sorbus aria, Sorbus aucuparia, Taxus baccata, Ulmus glabra, Sambucus nigra, Ruscus aculeatus, Ligustrum vulgare, Carex remota, Carex sylvatica, Luzula pilosa, Mnium hornum, Dicranella heteromalla, Dicranum scoparium, Fissidens taxifolius, Brachythecium rutabulum, Hypnum cupressiforme, Isothecium myosuroides, Kindbergia praelonga, Plagiothecium denticulatum, Plagiothecium undulatum, Pseudotaxiphyllum elegans, Thuidium tamariscinum, Atrichum undulatum, Polytrichum commune, Polytrichum formosum, Arum maculatum, Lonicera periclymenum, Moehringia trinervia, Stellaria media agg., Euphorbia amygdaloides, Mercurialis perennis, Hypericum pulchrum, Lamiastrum galeobdolon, Hyacinthoides non-scripta, Circaea lutetiana, Epipactis helleborine, Oxalis acetosella, Digitalis purpurea, Anemone nemorosa, Ranunculus repens, Geum urbanum, Rubus fruticosus agg., Rubus idaeus, Galium odoratum, Galium verum, Urtica dioica, Viola riviniana, Agrostis capillaris, Agrostis stolonifera, Brachypodium sylvaticum, Bromopsis ramosa, Dactylis glomerata, Deschampsia cespitosa, Deschampsia flexuosa, Holcus lanatus, Holcus mollis, Melica uniflora, Milium effusum, Poa nemoralis, Poa trivialis, Schedonorus giganteus, Pteridium aquilinum, Dryopteris dilatata, Dryopteris filix-mas

W15 — Beech wood, Wood pasture

Fagus sylvatica-Deschampsia flexuosa woodland

Hedera helix, Acer pseudoplatanus, Ilex aquifolium, Betula pendula, Betula pubescens, Fagus sylvatica, Quercus hybrid, Quercus petraea, Quercus robur, Pinus sylvestris, Sorbus aucuparia, Taxus baccata, Calluna vulgaris, Rhododendron ponticum, Vaccinium myrtillus, Ruscus aculeatus, Carex pilulifera, Luzula pilosa, Luzula sylvatica, Mnium hornum, Orthodontium lineare, Dicranella heteromalla, Dicranum scoparium, Leucobryum glaucum, Hylocomium splendens, Hypnum cupressiforme, Kindbergia praelonga, Plagiothecium undulatum, Pseudotaxiphyllum elegans, Rhytidiadelphus loreus, Thuidium tamariscinum, Polytrichum formosum, Tetraphis pellucida, Calypogeia fissa, Diplophyllum albicans, Gymnocolea inflata, Lepidozia reptans, Cladonia fimbriata, Cladonia squamules, Lonicera periclymenum, Hyacinthoides non-scripta, Chamerion angustifolium, Oxalis acetosella, Rubus fruticosus agg., Melanpyrum pratense, Agrostis capillaris, Deschampsia flexuosa, Holcus lanatus, Holcus mollis, Molinia caerulea, Poa nemoralis, Blechnum spicant, Pteridium aquilinum, Dryopteris dilatata

W16 — Atlantic oak wood, Lowland dry oak and birch wood, Wood pasture

Quercus spp.-Betula spp.-Deschampsia flexuosa woodland

Hedera helix, Acer pseudoplatanus, Ilex aquifolium, Betula pendula, Betula pubescens, Corylus avellana, Fagus sylvatica, Castanea sativa, Quercus hybrid, Quercus petraea, Quercus robur, Pinus sylvestris, Sorbus aria, Sorbus aucuparia, Populus tremula, Calluna vulgaris, Erica cinerea, Rhododendron ponticum, Vaccinium myrtillus, Ulex europaeus, Ulex gallii, Frangula alnus, Crataegus monogyna, Luzula sylvatica, Mnium hornum, Orthodontium lineare, Pohlia nutans, Campylopus flexuosus, Campylopus pyriformis, Dicranella heteromalla, Dicranodontium denudatum, Dicranoweisia cirrata, Dicranum fuscescens, Dicranum scoparium, Leucobryum glaucum, Hypnum cupressiforme, Hypnum jutlandicum, Isothecium myosuroides, Kindbergia praelonga, Plagiothecium denticulatum, Plagiothecium nemorale, Plagiothecium succulentum, Plagiothecium undulatum, Pleurozium schreberi, Pseudoscleropodium purum, Pseudotaxiphyllum elegans, Rhytidiadelphus loreus, Rhytidiadelphus triquetrus, Thuidium tamariscinum, Tetraphis pellucida, Barbilophozia attenuata, Barbilophozia floerkei, Calypogeia fissa, Calypogeia trichomanis, Cephalozia bicuspidata, Gymnocolea inflata, Lepidozia reptans, Lophocolea bidentata, Lophocolea heterophylla, Cladonia coccifera, Cladonia coniocraea, Cladonia fimbriata, Cladonia polydactyla, Cladonia squamosa, Solidago virgaurea, Lonicera periclymenum, Corydalis claviculata, Teucrium scorodonia, Hyacinthoides non-scripta, Chamerion angustifolium, Oxalis acetosella, Digitalis purpurea, Rumex acetosella, Potentilla erecta, Rubus fruticosus agg., Galium saxatile, Agrostis capillaris, Agrostis stolonifera, Anthoxanthum odoratum, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Festuca rubra, Holcus lanatus, Holcus mollis, Molinia caerulea, Blechnum spicant, Pteridium aquilinum, Dryopteris dilatata

W17 — Altantic hazel wood, Atlantic oak wood, Caledonian forest, Upland birch wood, Wood pasture

Quercus petraea-Betula pubescens-Dicranum majus woodland

Hedera helix, Salix caprea, Acer pseudoplatanus, Ilex aquifolium, Betula pendula, Betula pubescens, Corylus avellana, Fagus sylvatica, Quercus hybrid, Quercus petraea, Quercus robur, Fraxinus excelsior, Sorbus aucuparia, Calluna vulgaris, Erica cinerea, Erica tetralix, Juniperus communis, Rhododendron ponticum, Vaccinium myrtillus, Crataegus monogyna, Carex pilulifera, Luzula multiflora, Luzula pilosa, Luzula sylvatica, Andreaea rupestris, Mnium hornum, Rhizomnium punctatum, Campylopus flexuosus, Dicranella heteromalla, Dicranodontium denudatum, Dicranum fuscescens, Dicranum majus, Dicranum scoparium, Dicranum scottianum, Leucobryum glaucum, Ptychomitrium polyphyllum, Racomitrium fasciculare, Racomitrium heterostichum, Racomitrium lanuginosum, Eurhynchium striatum, Heterocladium heteropterum, Hylocomiastrum umbratum, Hylocomium armoricum, Hylocomium splendens, Hypnum callichroum, Hypnum cupressiforme, Hypnum jutlandicum, Isothecium myosuroides, Kindbergia praelonga, Loeskeobryum brevirostre, Plagiothecium denticulatum, Plagiothecium undulatum, Pleurozium schreberi, Pseudoscleropodium purum, Pseudotaxiphyllum elegans, Ptilium crista-castrensis, Rhytidiadelphus loreus, Rhytidiadelphus squarrosus, Rhytidiadelphus triquetrus, Sematophyllum demissum, Sematophyllum micans, Thuidium delicatulum, Thuidium tamariscinum, Polytrichum commune, Polytrichum formosum, Sphagnum capillifolium, Sphagnum fimbriatum, Sphagnum palustre, Sphagnum russowii, Sphagnum subnitens, Tetraphis pellucida, Adelanthus decipiens, Anastrepta orcadensis, Barbilophozia attenuata, Barbilophozia floerkei, Bazzania tricrenata, Bazzania trilobata, Blepharostoma trichophyllum, Calypogeia fissa, Calypogeia muelleriana, Cephalozia bicuspidata, Diplophyllum albicans, Frullania tamarisci, Fuscocephaloziopsis lunulifolia, Harpanthus scutatus, Jamesoniella autumnalis, Jungermannia gracillima, Lepidozia pearsonii, Lepidozia reptans, Lophocolea bidentata, Lophocolea heterophylla, Lophozia ventricosa, Marsupella emarginata, Microlejeunea ulicina, Mylia taylorii, Nowellia curvifolia, Plagiochila asplenioides, Plagiochila atlantica, Plagiochila killarniensis, Plagiochila porelloides, Plagiochila punctata, Plagiochila spinulosa, Ptilidium ciliare, Saccogyna viticulosa, Scapania gracilis, Scapania nemorea, Scapania umbrosa, Tritomaria exsectiformis, Tritomaria quinquedentata, Cladonia arbuscula, Cladonia chlorophaea, Cladonia digitata, Cladonia furcata, Cladonia impexa, Cladonia polydactyla, Cladonia squamosa, Cladonia squamules, Cladonia subcervicornis, Parmelia saxatilis, Sphaerophorus fragilis, Sphaerophorus globosus, Solidago virgaurea, Lonicera periclymenum, Succisa pratensis, Cornus suecica, Hypericum pulchrum, Teucrium scorodonia, Hyacinthoides non-scripta, Goodyera repens, Oxalis acetosella, Digitalis purpurea, Rumex acetosa, Lysimachia nemorum, Trientalis europaea, Anemone nemorosa, Potentilla erecta, Rubus fruticosus agg., Rubus idaeus, Galium saxatile, Melanpyrum pratense, Viola riviniana, Agrostis capillaris, Agrostis stolonifera, Agrostis vinealis, Anthoxanthum odoratum, Dactylis glomerata, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Holcus lanatus, Holcus mollis, Molinia caerulea, Poa nemoralis, Poa pratensis, Blechnum spicant, Pteridium aquilinum, Dryopteris aemula, Dryopteris borreri, Dryopteris dilatata, Dryopteris filix-mas, Hymenophyllum tunbrigense, Hymenophyllum wilsonii, Polypodium vulgare, Thelypteris limbosperma, Athyrium filix-femina

W20 — Montane scrub

Salix lapponum-Luzula sylvatica scrub

Salix arbuscula, Salix cinerea, Salix lapponum, Salix myrsinites, Salix phylicifolia, Salix reticulata, Empetrum nigrum, Calluna vulgaris, Salix lanata, Vaccinium myrtillus, Vaccinium uliginosum, Carex atrata, Carex bigelowii, Carex binervis, Carex flacca, Luzula multiflora, Luzula sylvatica, Huperzia selago, Selaginella selaginoides, Aulacomnium palustre, Mnium hornum, Philonotis fontana, Plagiomnium undulatum, Rhizomnium punctatum, Dicranum bonjeanii, Dicranum majus, Dicranum scoparium, Fissidens adianthoides, Racomitrium lanuginosum, Calliergonell cuspidatum, Ctenidium molluscum, Hylocomium splendens, Hypnum callichroum, Hypnum cupressiforme, Palustriella commutatum, Plagiothecium denticulatum, Plagiothecium undulatum, Pleurozium schreberi, Ptilium crista-castrensis, Rhytidiadelphus loreus, Rhytidiadelphus squarrosus, Rhytidiadelphus triquetrus, Sanionia uncinata, Scorpidium revolvens, Thuidium tamariscinum, Atrichum undulatum, Polytrichastrum alpinum, Sphagnum capillifolium, Sphagnum fallax, Sphagnum girgensohnii, Sphagnum palustre, Sphagnum russowii, Sphagnum squarrosum, Sphagnum subnitens, Sphagnum teres, Barbilophozia barbata, Chiloscyphus polyanthos, Diplophyllum albicans, Lophocolea bidentata, Plagiochila porelloides, Ptilidium ciliare, Scapania undulata, Cladonia squamosa, Peltigera canina, Angelica sylvestris, Caltha palustris, Hieraclium spp., Saussurea alpina, Solidago virgaurea, Campanula rotundifolia, Succisa pratensis, Valeriana officinalis, Rhodiola rosea, Vaccinium vitis-idaea, Thymus praecox, Epilobium anagallidifolium, Dactylorhiza viridis, Euphrasia officinalis, Rhinanthus minor, Oxalis acetosella, Oxyria digyna, Polygonum viviparum, Rumex acetosa, Anemone nemorosa, Ranunculus acris, Thalictrum alpinum, Alchemilla alpina, Alchemilla glabra, Filipendula ulmaria, Geum rivale, Galium boreale, Galium saxatile, Saxifraga aizoides, Saxifraga hypnoides, Saxifraga oppositifolia, Saxifraga stellaris, Viola riviniana, Agrostis canina, Agrostis capillaris, Anthoxanthum odoratum, Deschampsia cespitosa, Deschampsia flexuosa, Festuca ovina, Festuca rubra, Nardus stricta, Dryopteris dilatata, Phegopteris connectilis

W21 — Hedgerow scrub, Mixed scrub

Crataegus monogyna-Hedera helix scrub

Hedera helix, Salix cinerea, Acer pseudoplatanus, Ilex aquifolium, Betula pendula, Corylus avellana, Fagus sylvatica, Quercus robur, Fraxinus excelsior, Malus sylvestris, Sorbus aria, Taxus baccata, Ulmus glabra, Ulmus procera, Sambucus nigra, Viburnum lantana, Euonymus europaeus, Cornus sanguinea, Juniperus communis, Ligustrum vulgare, Clematis vitalba, Rhamnus cathartica, Crataegus laevigata, Crataegus monogyna, Prunus spinosa, Juncus effusus, Plagiomnium undulatum, Brachythecium rutabulum, Kindbergia praelonga, Rhynchostegium confertum, Tamus communis, Heracleum sphondylium, Torilis japonica, Arum maculatum, Arctium minus, Cirsium arvense, Lonicera periclymenum, Moehringia trinervia, Silene dioica, Stellaria graminea, Stellaria holostea, Stellaria media agg., Calystegia sepium, Mercurialis perennis, Geranium robertianum, Iris foetidissima, Glechoma hederacea, Lamium purpureum, Origanum vulgare, Stachys sylvatica, Teucrium scorodonia, Allium ursinum, Hyacinthoides non-scripta, Circaea lutetiana, Rumex sanguineus, Anemone nemorosa, Ranunculus repens, Fragaria vesca, Geum urbanum, Potentilla sterilis, Poterium sanguisorba, Rosa arvensis, Rosa canina, Rubia peregrina, Rubus fruticosus agg., Galium aparine, Urtica dioica, Viola riviniana, Anisantha sterilis, Arrhenatherum elatius, Brachypodium pinnatum, Brachypodium sylvaticum, Bromopsis erecta, Dactylis glomerata, Elytrigia repens, Holcus lanatus, Holcus mollis, Poa trivialis, Asplenium scolopendrium, Pteridium aquilinum, Dryopteris filix-mas

W22 — Hedgerow scrub, Mixed scrub

Prunus spinosa-Rubus fruticosus scrub

Hedera helix, Acer campestre, Corylus avellana, Ulex europaeus, Prunus spinosa, Plagiomnium undulatum, Brachythecium rutabulum, Eurhynchium striatum, Kindbergia praelonga, Thuidium tamariscinum, Atrichum undulatum, Lophocolea bidentata, Achillea millefolium, Cardamine flexuosa, Lonicera periclymenum, Moehringia trinervia, Silene dioica, Silene uniflora, Stellaria media agg., Mercurialis perennis, Geranium robertianum, Stachys sylvatica, Teucrium scorodonia, Hyacinthoides non-scripta, Oxalis acetosella, Digitalis purpurea, Plantago lanceolata, Veronica chamaedrys, Armeria maritima, Rumex acetosa, Filipendula ulmaria, Geum urbanum, Rosa canina, Rubus fruticosus agg., Rubus idaeus, Galium aparine, Urtica dioica, Viola riviniana, Agrostis capillaris, Brachypodium sylvaticum, Dactylis glomerata, Festuca rubra, Holcus lanatus, Holcus mollis, Poa trivialis, Asplenium scolopendrium, Pteridium aquilinum, Dryopteris borreri, Dryopteris filix-mas

W25 — Bracken scrub

Pteridium aquilinum-Rubus fruticosus underscrub

Hedera helix, Acer pseudoplatanus, Fagus sylvatica, Quercus robur, Fraxinus excelsior, Sambucus nigra, Ulex europaeus, Crataegus monogyna, Prunus spinosa, Luzula multiflora, Mnium hornum, Brachythecium rutabulum, Kindbergia praelonga, Angelica sylvestris, Conopodium majus, Heracleum sphondylium, Arum maculatum, Cirsium arvense, Lonicera periclymenum, Silene dioica, Stellaria holostea, Mercurialis perennis, Geranium robertianum, Glechoma hederacea, Teucrium scorodonia, Hyacinthoides non-scripta, Chamerion angustifolium, Circaea lutetiana, Digitalis purpurea, Rumex acetosa, Anemone nemorosa, Ficaria verna, Potentilla erecta, Rosa arvensis, Rosa canina, Rubus fruticosus agg., Rubus idaeus, Galium aparine, Galium mollugo sens.lat., Galium saxatile, Urtica dioica, Viola riviniana, Agrostis capillaris, Anthoxanthum odoratum, Arrhenatherum elatius, Dactylis glomerata, Festuca rubra, Holcus lanatus, Holcus mollis, Poa trivialis, Pteridium aquilinum, Dryopteris dilatata, Dryopteris filix-mas